Phthinosuchus Temporal range: Middle Permian, | |
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Life restoration of Phthinosuchus discors | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Synapsida |
Clade: | Therapsida |
Suborder: | † Dinocephalia (?) |
Family: | † Phthinosuchidae Yefremov, 1954 |
Genus: | † Phthinosuchus Yefremov, 1954 |
Type species | |
†Phthinosuchus discors Yefremov, 1954 |
Phthinosuchus is an extinct genus of therapsids from the Middle Permian of Russia. Phthinosuchus is the sole member of the family Phthinosuchidae. Phthinosuchus may have been one of the most primitive therapsids, meaning that its ancestors may have branched off early from the main therapsid line.
Phthinosuchus was named in 1954 by Ivan Yefremov. It is currently only known from the back of the skull, as the front of the skull was lost after description.[ clarification needed ] Two species, P. discors and P. horissiaki, have been described. [1]
Phthinosuchus was 1.5 m (5 ft) long with a 20 cm skull, and looked much like the Sphenacodontids, such as Dimetrodon and Sphenacodon . Its temporal fenestrae were larger than those of the Sphenacodontids. [2] [3] Its jaw was slender, unlike other predatory therapsids, [1] but like the other early therapsids, it was probably sprawling and carnivorous.
Phthinosuchus is a member of the suborder Phthinosuchia, which was made specifically for Phthinosuchus given that it did not fit into any other suborder. [1]
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typically toothless beak, unique amongst all synapsids. Dicynodonts first appeared in Southern Pangaea during the mid-Permian, ca. 270–260 million years ago, and became globally distributed and the dominant herbivorous animals in the Late Permian, ca. 260–252 Mya. They were devastated by the end-Permian Extinction that wiped out most other therapsids ca. 252 Mya. They rebounded during the Triassic but died out towards the end of that period. They were the most successful and diverse of the non-mammalian therapsids, with over 70 genera known, varying from rat-sized burrowers to elephant-sized browsers.
Dinocephalians are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania.
Prenocephale is a genus of small pachycephalosaurid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was similar in many ways to its close relative, Homalocephale.
Eupelycosauria is a large clade of animals characterized by the unique shape of their skull, encompassing all mammals and their closest extinct relatives. They first appeared 308 million years ago during the Early Pennsylvanian epoch, with the fossils of Echinerpeton and perhaps an even earlier genus, Protoclepsydrops, representing just one of the many stages in the evolution of mammals, in contrast to their earlier amniote ancestors.
Biarmosuchia is an extinct clade of non-mammalian synapsids from the Permian. Biarmosuchians are the most basal group of the therapsids. They were moderately-sized, lightly-built carnivores, intermediate in form between basal sphenacodont "pelycosaurs" and more advanced therapsids. Biarmosuchians were rare components of Permian ecosystems, and the majority of species belong to the clade Burnetiamorpha, which are characterized by elaborate cranial ornamentation.
The theriodonts are a major group of therapsids which appeared during the Middle Permian and which includes the gorgonopsians and the eutheriodonts, itself including the therocephalians and the cynodonts.
Sphenacodon is an extinct genus of synapsid that lived from about 300 to about 280 million years ago (Ma) during the Late Carboniferous and Early Permian periods. Like the closely related Dimetrodon, Sphenacodon was a carnivorous member of the Eupelycosauria family Sphenacodontidae. However, Sphenacodon had a low crest along its back, formed from blade-like bones on its vertebrae instead of the tall dorsal sail found in Dimetrodon. Fossils of Sphenacodon are known from New Mexico and the Utah–Arizona border region in North America.
Titanosuchus ferox is an extinct species of dinocephalian therapsids that lived in the Middle Permian epoch in South Africa.
Lycaenops ("wolf-face") is a genus of carnivorous therapsids. It lived during the Middle Permian to the early Late Permian, about 260 mya, in what is now South Africa.
Cainotherium is an extinct genus of rabbit-sized prehistoric even-toed ungulates. These herbivores lived in Europe from the Eocene until the early Miocene. The skeletal anatomy of these hare-like animals suggest they, along with other members of Cainotheriidae, belong to the artiodactyl suborder Tylopoda, together with oreodonts and modern camelids. Species had cloven hooves, similar to those of bovids or deer, although the shape and length of the limbs suggests that the living animals moved by leaping, like a rabbit. The shape of the teeth also suggests a rabbit-like diet of nibbled vegetation, while the size of the auditory bulla and shape of the brain suggest that it would have had good senses of hearing and smell.
Peltobatrachus is an extinct genus of temnospondyl amphibian from the late Permian period of Tanzania. The sole species, Peltobatrachus pustulatus, is also the sole member of the family Peltobatrachidae.
Araeoscelis is an extinct genus of reptile, and one of the earliest diapsids. Fossils have been found in the Nocona, Arroyo and Waggoner Ranch Formations in Texas, dating to the Early Permian. Two species have been described, A. casei and A. gracilis.
Hovasaurus is an extinct genus of diapsid reptile belonging to the order Eosuchia. It lived in what is now Madagascar during the Late Permian and Early Triassic, being a survivor of the Permian–Triassic extinction event and the paleontologically youngest member of the Tangasauridae. Fossils have been found in the Permian Lower and Triassic Middle Sakamena Formations of the Sakamena Group, where it is amongst the commonest fossils. Its morphology suggests an aquatic ecology.
Galechirus is an extinct genus of anomodont therapsids. It was about 30 cm (1 ft) long.
Pachyrukhos is an extinct genus of hegetotheriid notoungulate from the Early to Middle Miocene of Argentina and Chile. Fossils of this genus have been found in the Collón Curá, Sarmiento and Santa Cruz Formations of Argentina and the Río Frías Formation of Chile.
Adinotherium is an extinct genus of Toxodontidae, large bodied hoofed ungulates which inhabited South America during the Middle to Late Miocene, from 17.5 to 6.8 Ma and existed for approximately 10.7 million years, Santacrucian to Huayquerian in the South American land mammal ages (SALMA). Fossils of Adinotherium have been found in the Santa Cruz and Ituzaingó Formations of Argentina and the Chucal and Río Frías Formations of Chile.
Pantydraco was a genus of basal sauropodomorph dinosaur from the Late Triassic of the United Kingdom (Wales). It is based on a partial juvenile skeleton once thought to belong to Thecodontosaurus. Only one valid species of Pantydraco is recognised: P. caducus.
Microurania is an extinct genus of therapsids from the Middle Permian first named and described by Mikhaïl Ivakhnenko. It is known from a single partial skull found in the region of Orenburg, Russia. According to Kammerer, 2011, it likely represents the remains of a juvenile dinocephalian.
Herpetoskylax is an extinct genus of biarmosuchians which existed in South Africa. The type species is Herpetoskylax hopsoni. It lived in the Late Permian Period.
Phthinosuchia is an extinct group of therapsids including two poorly known species, Phthinosuchus discors and Phthinosaurus borrisiaki, from the Middle Permian of Russia. Phthinthosuchus is known a partial crushed skull and Phthinosaurus is known from an isolated lower jaw. The two species have traditionally been grouped together based on their shared primitive characteristics, but more recent studies have proposed that they are more distantly related. Phthinosuchus is either a carnivorous gorgonopsian relative or an anteosaurian dinocephalian while Phthinosaurus is either a herbivorous rhopalodont dinocephalian or a therocephalian.