Eomanis

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Eomanis
Temporal range: 47.8–46.3  Ma
O
S
D
C
P
T
J
K
Pg
N
Middle Eocene
Eomanis waldi.jpg
Fossil of Eomanis waldi
Eomanis NT.jpg
Life reconstruction of
Eomanis waldi
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Pholidota
Suborder: Eupholidota
Superfamily: Eomanoidea
Gaudin, 2009 [1]
Family: Eomanidae
Storch, 2003 [2]
Genus: Eomanis
Storch, 1978 [3]
Type species
Eomanis waldi
Storch, 1978

Eomanis ("dawn pangolin") is the earliest known true (and scaled) pangolin from extinct family Eomanidae (and extinct superfamily Eomanoidea) within suborder Eupholidota. It lived during the Eocene in Europe. [4] Eomanis fossils found in the Messel Pit in Germany are very similar in size and anatomy to living pangolins of the genus Manis , indicating that pangolins have remained largely unchanged in morphology and behavior for 50 million years. However, unlike modern pangolins, its tail and legs did not bear scales.[ citation needed ] According to the stomach contents of the excellently preserved Messel specimens, Eomanis’ diet consisted of both insects and plants.

Contents

Another early mammal discovered in the Messel Pit that was similar in anatomy and likely also had an insectivorous diet was Eurotamandua . [5] Despite its name, Eurotamandua was almost certainly not a xenarthran because it lacked the characteristic xenarthran joints present in all living and extinct xenarthrans. In addition, xenarthrans existed exclusively in South America for almost the entire Cenozoic era. They spread to North America after the Panama land bridge formed 3 million years ago, but never reached Eurasia or Africa. Eurotamandua was likely also an early pangolin, possibly the ancestor of Eomanis.

Phylogeny

Phylogenetic position of genus Eomanis within order Pholidota. [1] [6]

  Ferae  

Pan-Carnivora Dogs, jackals, wolves, and foxes (Plate XI).jpg

  Pholidotamorpha  

Palaeanodonta Metacheiromys DB152-2.jpg

  Pholidota  

Euromanis

 ? 

Pholidota sp. (BC 16’08)

Eurotamanduidae

  Eupholidota  

Manoidea Pangolin Hardwicke (white background).jpg

 Eomanoidea 
 Eomanidae 
 Eomanis 

Eomanis waldi

 sensu stricto 
 (Pholidota [sensu lato]) 

See also

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References

  1. 1 2 Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. Heidelberg, Germany: Springer Science+Business Media. 16 (4): 235–305. doi:10.1007/s10914-009-9119-9. S2CID   1773698. Archived from the original (PDF) on 2015-09-25. Retrieved 2020-11-16.
  2. Gerhard Storch (2003) "Fossil Old World "edentates" (Mammalia)." Senckenbergiana biologica 83 (1), pp. 51-60
  3. Gerhard Storch (1978) "Eomanis waldi, a pangolin from the Middle Eocene of the "Pit Messel" near Darmstadt (Mammalia: Pholidota)." Senckenbergiana lethaea 59 (4/6), pp. 503-529
  4. Cox, Barry; Savage, R. J. G.; Gardiner, Brian; Dixon, Dougal (1988). "Insectivores and creodonts". Macmillan Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. Macmillan London Limited. p. 213. ISBN   0-333-48699-4.
  5. Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 209. ISBN   1-84028-152-9.
  6. Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. Bibcode:2012JVPal..32..983K. doi:10.1080/02724634.2012.694319. ISSN   0272-4634. S2CID   86059673.