Necromanis

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Necromanis
Temporal range: 28.8–12.75  Ma
O
S
D
C
P
T
J
K
Pg
N
Middle Oligocene to Middle Miocene
Comparison Sansanosmilus palmidens - Necromanis franconica.jpg
N. franconica and Sansanosmilus
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Pholidota
Suborder: Eupholidota
Superfamily: Manoidea
Genus: Necromanis
Filhol, 1894
Type species
Necromanis franconica
Quenstedt, 1886
Species
  • N. franconica(Quenstedt, 1885) [1]
  • N. parva(Koenigswald, 1969) [2]
  • N. quercyi(Filhol, 1894) [3]
Synonyms
synonyms of genus:
  • Galliaetatus(Ameghino, 1905)
  • Leptomanis(Filhol, 1894)
  • Necrodasypus(Filhol, 1894)
  • Teutomanis(Ameghino, 1905)
synonyms of species:
  • N. franconica:
    • Galliaetatus schlosseri(Ameghino, 1905)
    • Lutra franconica(Quenstedt, 1885)
    • Teutomanis franconica
    • Teutomanis quenstedti(Ameghino, 1905)
  • N. quercyi:
    • Leptomanis edwardsi(Filhol, 1894)
    • Necromanis edwardsi(Koenigswald & Martin, 1990) [4]

Necromanis ("extinct pangolin") is an extinct genus of pangolin from superfamily Manoidea. It lived from the middle Oligocene to middle Miocene in Europe. It was originally placed within family Manidae, but was eventually removed from it as more fossil pholidotids from outside that family were found and studied more extensively (i.e., with the discovery and study of Eomanis and Patriomanis ). [5] Currently, Necromanis is placed as incertae sedis within the pholidotid superfamily Manoidea, together with the families Manidae and Patriomanidae. [6]

Contents

N. quercyi was originally placed within Teutomanis by Ameghino in 1905, but was later subsumed into Necromanis. A new fossil humerus attributed to N. franconica from Quercy, France lead researchers to reaffirm Teutomanis quercyi's status as distinct from Necromanis. [7]

Phylogeny

Phylogenetic position of genus Necromanis within superfamily Manoidea. [6] [8]

  Ferae  

Pan-Carnivora Dogs, jackals, wolves, and foxes (Plate XI).jpg

  Pholidotamorpha  

Palaeanodonta Metacheiromys DB152-2.jpg

  Pholidota  

Euromanis

 ? 

Pholidota sp. (BC 16’08)

Eurotamanduidae

  Eupholidota  

Eomanoidea

  Manoidea  

Manidae Pangolin Hardwicke (white background).jpg

Patriomanidae

 Necromanis 

Necromanis franconica

Necromanis parva

Necromanis quercyi

 ? 
 sensu stricto 
 (Pholidota [sensu lato]) 

See also

Related Research Articles

<i>Manis</i> Genus of mammals

Manis ("spirit") is a genus of South Asian and East Asian pangolins, the Asiatic pangolins, from subfamily Maninae, within family Manidae.

<span class="mw-page-title-main">Manidae</span> Family of pangolins

Manidae ("spirits") is the only extant family of pangolins from superfamily Manoidea. This family comprises three genera, as well as extinct Fayum pangolin.

<i>Eomanis</i> Extinct genus of mammals

Eomanis is the earliest known true pangolin from extinct family Eomanidae within suborder Eupholidota. It lived during the Eocene in Europe. Eomanis fossils found in the Messel Pit in Germany are very similar in size and anatomy to living pangolins of the genus Manis, indicating that pangolins have remained largely unchanged in morphology and behavior for 50 million years. However, unlike modern pangolins, its tail and legs did not bear scales. According to the stomach contents of the excellently preserved Messel specimens, Eomanis’ diet consisted of both insects and plants.

<i>Metacheiromys</i> Extinct genus of mammal

Metacheiromys is an extinct genus of placental mammals from extinct paraphyletic subfamily Metacheiromyinae within extinct paraphyletic family Metacheiromyidae in extinct order Palaeanodonta, that lived in North America from the early to middle Eocene.

<i>Eurotamandua</i> Extinct genus of pangolins

Eurotamandua is an extinct genus of mammal from extinct family Eurotamanduidae that lived during the middle Eocene.

<i>Ernanodon</i> Extinct genus of mammal

Ernanodon is an extinct genus of placental mammal from extinct family Ernanodontidae within extinct order Palaeanodonta, that lived from the middle to late Paleocene in China and Mongolia.

<span class="mw-page-title-main">Epoicotheriidae</span> Extinct family of mammals

Epoicotheriidae is an extinct paraphyletic family of insectivorous placental mammals within extinct order Palaeanodonta, that lived in North America, Asia and Europe from the late Paleocene to early Oligocene. Epoicotheriids were fossorial mammals. Late Eocene/early Oligocene genera were highly specialized animals that were convergent with the talpids, golden moles and marsupial mole in the structure of their skulls and forelimbs, and would have had a similar lifestyle as subterranean burrowers.

Escavadodon is an extinct genus of pangolin-like myrmecophagous placental mammals of extinct monotypic family Escavadodontidae within extinct order Palaeanodonta, that lived in North America during the middle Paleocene. It contains a single species, Escavadodon zygus, recovered from the Nacimiento Formation of New Mexico.

<i>Phataginus</i> Genus of mammals

African tree pangolin (Phataginus) is a genus of African pangolins from subfamily small African pangolins (Phatagininae), within family Manidae. Its members are the more arboreal of the African pangolins.

<i>Smutsia</i> Genus of mammals

African ground pangolin is a genus of pangolins from subfamily Smutsiinae within family Manidae. It was formerly considered a subgenus of genus Manis. Its members are the more terrestrial of the African pangolins. In past, this genus was also present in Europe.

<i>Patriomanis</i> Genus of mammals

Patriomanis is an extinct genus of pangolin from extinct family Patriomanidae. It lived from the late Eocene to early Oligocene of North America and it currently represents the only pangolin known from the Western Hemisphere. The genus contains one species, P. americana, which is known from six specimens, mostly from the Chadronian White River Formation of Montana. It had long digits and a prehensile tail, suggesting that it was arboreal, and its jaw was capable of opening wider than modern pangolins. Its ears and the hair between its scales were also longer than modern pangolins.

<span class="mw-page-title-main">Patriomanidae</span> Extinct family of pangolins

Patriomanidae is an extinct family of pangolins from superfamily Manoidea that includes two extinct genera Patriomanis and Cryptomanis.

<span class="mw-page-title-main">Pholidotamorpha</span> Clade of mammals

Pholidotamorpha is a clade of placental mammals from mirorder Ferae that includes the order Pholidota and extinct order Palaeanodonta.

<i>Euromanis</i> Genus of mammals

Euromanis is one of the earliest known pangolin genera. It lived during the middle Eocene in Europe. Euromanis fossils found in the Messel Pit in Germany. Unlike modern pangolins, it did not bear scales on its body.

<span class="mw-page-title-main">Eupholidota</span> Suborder of pangolins

Eupholidota is a suborder of pangolins that includes two superfamilies: extant Manoidea and extinct Eomanoidea.

<span class="mw-page-title-main">Manoidea</span> Superfamily of pangolins

Manoidea ("spirits") is a superfamily of pangolins from suborder Eupholidota that includes extant family Manidae, extinct family Patriomanidae and extinct genus Necromanis.

<i>Xenocranium</i> Genus of prehistoric burrowing mammals

Xenocranium is an extinct monotypic genus of placental mammal from extinct paraphyletic subfamily Epoicotheriinae within extinct paraphyletic family Epoicotheriidae in extinct order Palaeanodonta, that lived in North America during the late Eocene.

Smutsia olteniensis is an extinct species of pangolins from genus Smutsia of subfamily Smutsiinae within family Manidae. Fossilized remains of the species were found in Romania, providing evidence regarding the existence of pangolins in Europe during the Plio-Pleistocene period.

<span class="mw-page-title-main">Epoicotheriinae</span> Extinct subfamily of mammals

Epoicotheriinae is an extinct paraphyletic subfamily of insectivorous placental mammals within extinct paraphyletic family Epoicotheriidae in extinct order Palaeanodonta, that lived in North America and Europe from the early Eocene to early Oligocene. Epoicotheriins were fossorial mammals. Late Eocene/early Oligocene genera were highly specialized animals that were convergent with the talpids, golden moles and marsupial mole in the structure of their skulls and forelimbs, and would have had a similar lifestyle as subterranean burrowers.

Cryptomanis is an extinct genus of pangolin from extinct family Patriomanidae. The genus is only known from the holotype specimen from middle Eocene deposits from Inner Mongolia, China. The holotype, AMNH 26140, was for years labeled as an unnamed pangolin in the fossil collection for decades. The remains comprise of an incomplete postcranial specimen, with nearly complete hind limbs, an incomplete pelvis, a complete lumbar series from an incomplete vertebral column, numerous fragmentary remains of rib and sternal bones, a fragmentary scapula, and partially preserved forelimbs. Based on the femural length Cryptomanis was comparable in size to the modern Sunda pangolin.

References

  1. Quenstedt, F. A. (1885.) "Handbuch der Petrefaktenkunde 4th ed." Verlag der H. Lauppschen Buchhandlung, Tübingen, 1239 p.
  2. Von Wighart, Koenigswald (1969.) "Die Maniden (Pholidota, Mamm.) des europäischen Tertiärs", München, Bayerische Staatssammlung für Paläontologie und Historische Geologie, 9, 61-71
  3. Filhol, M. H. (1894.) "Observations concernant quelques mammifères fossiles nouveaux du Quercy." Annales de Sciences Naturelles (Zoologie), 16, 129-150
  4. Wighart Von Koenigswald, T. Martin (1990.) A skeleton of Necromanis franconica, a pangolin (Pholidota, Mammalia) of the Aquitanian from Saulcet in the Allier Basin, France, Eclogae Geologicae Helvetiae 83(3):845-864
  5. Gaudin, Timothy J., Robert J. Emry, and Brandon Pogue. "A new genus and species of pangolin (Mammalia, Pholidota) from the late Eocene of Inner Mongolia, China." Journal of Vertebrate Paleontology 26.1 (2006): 146-159.
  6. 1 2 Gaudin, Timothy (2009). "The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis" (PDF). Journal of Mammalian Evolution. Heidelberg, Germany: Springer Science+Business Media. 16 (4): 235–305. doi:10.1007/s10914-009-9119-9. S2CID   1773698. Archived from the original (PDF) on 2015-09-25. Retrieved 2020-12-18.
  7. CROCHET, J. Y., HAUTIER, L., & LEHMANN, T. (2015). A pangolin (Manidae, Pholidota, Mammalia) from the French Quercy phosphorites (Pech du Fraysse, Saint-Projet, Tarn-et-Garonne, late Oligocene, MP 28). Palaeovertebrata, 39(2), e4.
  8. Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of Ernanodon (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis". Journal of Vertebrate Paleontology. 32 (5): 983–1001. Bibcode:2012JVPal..32..983K. doi:10.1080/02724634.2012.694319. ISSN   0272-4634. S2CID   86059673.