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A squid is a mollusc with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes. Like all other cephalopods, squid have a distinct head, bilateral symmetry, and a mantle. They are mainly soft-bodied, like octopuses, but have a small internal skeleton in the form of a rod-like gladius or pen, made of chitin.
A cephalopod is any member of the molluscan class Cephalopoda such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology.
The giant squid is a species of deep-ocean dwelling squid in the family Architeuthidae. It can grow to a tremendous size, offering an example of deep-sea gigantism: recent estimates put the maximum size at around 12–13 m (39–43 ft) for females and 10 m (33 ft) for males, from the posterior fins to the tip of the two long tentacles. The mantle of the giant squid is about 2 m long, and the length of the squid excluding its tentacles rarely exceeds 5 m (16 ft). Claims of specimens measuring 20 m (66 ft) or more have not been scientifically documented.
Subclass Coleoidea, or Dibranchiata, is the grouping of cephalopods containing all the various taxa popularly thought of as "soft-bodied" or "shell-less". Unlike its extant sister group, Nautiloidea, whose members have a rigid outer shell for protection, the coleoids have at most an internal cuttlebone, gladius, or shell that is used for buoyancy or support. Some species have lost their cuttlebone altogether, while in some it has been replaced by a chitinous support structure. A unique trait of the group is the ability to edit their own RNA.
Kronosaurus is a potentially dubious genus of extinct short-necked pliosaur. With an estimated length of 9 to 10.9 meters, it was among the largest pliosaurs, and is named after the leader of the Greek Titans, Kronos. It lived in the Early Cretaceous period. Fossil material has been recovered from the Toolebuc Formation and Wallumbilla Formations (Aptian) of Queensland and New South Wales in Australia.
Cephalopods vary enormously in size. The smallest are only about 1 centimetre (0.39 in) long and weigh less than 1 gram (0.035 oz) at maturity, while the largest—the giant and colossal squids—can exceed 10 metres (33 ft) in length and weigh close to half a tonne (1,100 lb), making them the largest living invertebrates. Living species range in mass more than three-billion-fold, or across nine orders of magnitude, from the lightest hatchlings to the heaviest adults. Certain cephalopod species are also noted for having individual body parts of exceptional size. The giant and colossal squids, for example, have the largest known eyes among living animals.
Tusoteuthis is a extinct genus of large enchoteuthine cephalopod that lived during the Cretaceous. Although often called a squid, it is now thought to be more closely related to modern octopuses. Examination of gladius remains has yielded an estimated mantle length close to or equal to that of the modern giant squid. Fossil remains have been found in parts of the one-time Western Interior Seaway of North America, including Late Cretaceous rocks in Kansas, Colorado, Wyoming, South Dakota, North Dakota, and Manitoba Province. One species, T. longa, is traditionally recognized. In 2020, due to poor preservation of holotype specimen, genus Tusoteuthis is likely to be invalid, and it is considered that later described species are better to include in genus Enchoteuthis instead.
Boletzkyida is a primitive order of teuthid coleoid cephalopod: the boletzkyids are thought to be the earliest forms of coleoid cephalopods, and appear to form a link between nautiloid orthocerids and more advanced coleoids. Boletzkyida was named and described by Bandel, Reitner, and Sturmer in 1983 (B.R.&S) from specimens found in the Lower Devonian black slate in Germany.
Belemnotheutis is an extinct coleoid cephalopod genus from the middle and upper Jurassic, related to but morphologically distinct from belemnites. Belemnotheutis fossils are some of the best preserved among coleoids. Remains of soft tissue are well-documented in some specimens, even down to microscopic muscle tissue. In 2008, a group of paleontologists even recovered viable ink from ink sacs found in several specimens.
The Allaru Formation, also known as the Allaru Mudstone, is a geological formation in Queensland, Australia, whose strata date back to the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.
Acanthoteuthis is a belemnite genus, a squid-like cephalopod with an internal shell from the Late Jurassic Epoch, related to modern coleoids.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Belemnitida is an extinct order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous. Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most to the tip: the tongue-shaped pro-ostracum, the conical phragmocone, and the pointy guard. The calcitic guard is the most common belemnite remain. Belemnites, in life, are thought to have had 10 hooked arms and a pair of fins on the guard. The chitinous hooks were usually no bigger than 5 mm (0.20 in), though a belemnite could have had between 100 and 800 hooks in total, using them to stab and hold onto prey.
The gladius, or pen, is a hard internal bodypart found in many cephalopods of the superorder Decapodiformes and in a single extant member of the Octopodiformes, the vampire squid. It is so named for its superficial resemblance to the Roman short sword of the same name, and is a vestige of the ancestral mollusc shell, which was external. The gladius is located dorsally within the mantle and usually extends for its entire length. Composed primarily of chitin, it lies within the shell sac, which is responsible for its secretion.
All extant cephalopods have a two-part beak, or rostrum, situated in the buccal mass and surrounded by the muscular head appendages. The dorsal (upper) mandible fits into the ventral (lower) mandible and together they function in a scissor-like fashion. The beak may also be referred to as the mandibles or jaws.
Cephalopod fins, sometimes known as wings, are paired flap-like locomotory appendages. They are found in ten-limbed cephalopods as well as in the eight-limbed cirrate octopuses and vampire squid. Many extinct cephalopod groups also possessed fins. Nautiluses and the more familiar incirrate octopuses lack swimming fins. An extreme development of the cephalopod fin is seen in the bigfin squid of the family Magnapinnidae.
The Northumberland Formation is a Late Cretaceous (?Campanian-?Maastrichtian)-aged geologic formation in Canada. It belongs to the larger Nanaimo Group. Indeterminate bird and pterosaur fossils have been recovered from the formation, as well as a potential gladius of Eromangateuthis. The Northumberland Formation has been known to science since at least 2008.
Boreopeltis is an extinct genus of plesioteuthidid cephalopod, with 4 known species.
Enchoteuthis is an extinct genus of large enchoteuthine cephalopod that lived during the Cretaceous. Although it and its relative Tusoteuthis are often compared to squid, both are now thought to be more closely related to modern octopuses. Examination of gladius remains initially yielded an estimated mantle length about 2 m based on specimen once described as Tusoteuthis longa, close to or equal to that of the modern giant squid, although reclassification of this genus as a muensterelloid results in a much shorter total length, about 3 m (10 ft). Three species are currently recognized as valid: E. melanae, E. tonii, and E. cobbani.
The Teudopseina is a clade of stem-octopods that first evolved in the Toarcian, considered the largest clade of gladius-bearing coleoids in the Mesozoic. Up to five families are known, among which the Trachyteuthidae, Trachyteuthis in particular, are the most abundant.
References
- 1 2 3 4 5 Fuchs, D. (2019). "Eromangateuthis n. gen., a new genus for a late Albian gladius-bearing giant octobrachian (Cephalopoda: Coleoidea)". Paleontological Contributions. 21: 1–3. doi: 10.17161/1808.29619 .
- 1 2 3 Wade, M. (1993). "New Kelaenida and Vampyromorpha: Cretaceous squid from Queensland". Memoirs of the Association of Australasian Palaeontologists. 15: 353–374.
- ↑ Fuchs, D.; Beard, G.; Tanabe, K.; Ross, R. (2007). "Coleoid cephalopods from the Late Cretaceous north eastern Pacific". Seventh International Symposium 'Cephalopods - Present & Past'. Abstracts Volume (PDF). Sapporo, Japan: Hokkaido University. p. 131.
- ↑ Fuchs, D.; Stinnesbeck, W. (2021). "Large-sized gladius-bearing octobrachians (coleoid cephalopods) in the Turonian plattenkalk of Vallecillo, Mexico". Cretaceous Research. 127: 104949. doi:10.1016/j.cretres.2021.104949.