Northumberland Formation

Last updated
Northumberland Formation
Stratigraphic range: Late Cretaceous,
?Campanian–Maastrichtian
Type Geological formation
Unit of Nanaimo Group
Underlies Spray Formation
Location
Coordinates 49°36′N124°42′W / 49.6°N 124.7°W / 49.6; -124.7
Approximate paleocoordinates 56°06′N94°24′W / 56.1°N 94.4°W / 56.1; -94.4
Country Canada

The Northumberland Formation is a Late Cretaceous (?Campanian-?Maastrichtian)-aged geologic formation in Canada. It belongs to the larger Nanaimo Group. Indeterminate bird and pterosaur fossils have been recovered from the formation, [1] as well as a potential gladius of Eromangateuthis . [2] An extensive diversity of shark teeth is known from the formation; many appear to be closely allied with modern deep-water shark taxa, suggesting a deep-water environment for the formation. [3] The most well-known exposures of the formation are on Hornby Island.

Contents

Vertebrate paleofauna

Cartilaginous fish

Source: [4]

Hybodontiformes

GenusSpeciesLocationAbundanceNotesImages
Meristodonoides M. sp.A hybodontid shark.

Hexanchiformes

GenusSpeciesLocationAbundanceNotesImages
Chlamydoselachus C. balliA chlamydoselachid shark, related to the modern frilled shark. Chlamydoselachus anguineus2.jpg
Rolfodon R. ludvigseniA chlamydoselachid shark.
R. cf. thompsoni
Dykeius D. garethiA very large chlamydoselachid shark.
Hexanchus H. microdonA cow shark related to the modern bluntnose sixgill shark. Hexanchus griseus santa rosa.jpg
Notidanodon N. pectinatusA cow shark. Hexanchidae species - Tooth.jpg
Xampylodon X. dentatusA large cow shark. Notidanodon.jpg
Protoheptranchias P. loweiA cow shark similar to the modern sharpnose sevengill shark.
Paraorthacodus P. rossiA paraorthacodontid shark. Paraorthacodus.jpg
Komoksodon K. kwutchakuthA komoksodontid shark.

Echinorhiniformes

GenusSpeciesLocationAbundanceNotesImages
Echinorhinus E. lapaoiA echinorhinid shark, related to the modern bramble shark. Echinorhinus brucus Mc Coy.jpg

Squaliformes

GenusSpeciesLocationAbundanceNotesImages
Eoetmopterus E. supercretaceusAn etmopterid shark similar to modern lantern sharks.
Squalus S. vondermarckiA dogfish shark, related to modern spurdogs. Squalus acanthias stellwagen.jpg
S. nicholsae
S. sp.
Centrosqualus C. mustardiA dogfish shark.
Protocentrophorus P. steviaeA dogfish shark.
Rhinoscymnus R. clarkiA sleeper shark, possibly included within Somniosus . Somniosus rostratus.png
Centroscymnus C. sp.A sleeper shark related to the modern Portuguese dogfish. Centroscymnus coelolepis norfanz.jpg
Squaliodalatias S. savoieiA dalatiid shark.
Hessinodon H. wardiA possibly dalatiid shark.

Pristiophoriformes

GenusSpeciesLocationAbundanceNotesImages
Pristiophorus P. smithiA sawshark. Pristiophorus japonicus cropped.jpg
P. pricei

Orectolobiformes

GenusSpeciesLocationAbundanceNotesImages
Plicatoscyllium P. cf. globidensA ginglymostomatid shark.
Hemiscyllium H. hermaniA bamboo shark. Hemiscyllium halmahera.jpg

Lamniformes

GenusSpeciesLocationAbundanceNotesImages
Carcharias C. domingueiA sand shark, related to the modern sand tiger shark. Grey Nurse Shark at Fish Rock Cave, NSW.jpg

Synechodontiformes

GenusSpeciesLocationAbundanceNotesImages
Synechodus S. derekiA palaeospinacid shark.

Carcharhiniformes

GenusSpeciesLocationAbundanceNotesImages
Scyliorhinidae indent.A catshark of uncertain affinities.
Florenceodon F. johnyiA florenceodontid shark.

Bony fish

GenusSpeciesLocationAbundanceNotesImages
Gwawinapterus G. beardiJaw.A saurodontid ichthyodectiform, initially identified as an istiodactylid pterosaur. [5]

Birds

GenusSpeciesLocationAbundanceNotesImages
Maaqwi [6] M. cascadensisCoracoid and wing bones.A large ornithuran, either a vegaviid or a procellariform. [7] Maaqwi.png
Enantiornithes indet. [8] An enanthiornithine avialan.

Pterosaurs

GenusSpeciesLocationAbundanceNotesImages
Azhdarchoidea indet. [1] A possibly azhdarchid pterosaur.

Squamates

GenusSpeciesLocationAbundanceNotesImages
Mosasauridae indet. [9] Tylosaurus spA mosasaur.

Invertebrate paleofauna

Molluscs

Cephalopods

Genus [10] SpeciesLocationAbundanceNotesImages
Baculites B. occidentalisA baculitid ammonite. Baculites fossil cropped.png
Fresvillia F. constrictaA baculitid ammonite.
Diplomoceras D. cylindraceumA diplomoceratid ammonite. Diplomoceras.jpg
D. cf. cylindraceum
Exiteloceras E. densicostatumA diplomoceratid ammonite. Exiteloceras jenneyi 1.jpg
E. bipunctatum
Phylloptychoceras P. horitaiA diplomoceratid ammonite.
Solenoceras S. exornatusA diplomoceratid ammonite.
S. cf. reesidei
Nostoceras N. adrotansA nostoceratid ammonite. Nostoceras species cropped.jpg
N. hornbyensis
N. aff. pauper
Enchoteuthis [2] E. sp.A muensterellid octopodiform. Enchoteuthis reconstruction.png
Eromangateuthis [2] E. soniae?A plesioteuthid octopodiform.
Cyrtobelus [11] C. hornbyenseA groenlandibelid spirulid. [2]
Actinosepia [2] A. canadensisAn actinosepiid vampyromorphid. [12]
Cirroteuthidae indet. [2] Gladius.A cirroteuthid octopus.

Gastropods

GenusSpeciesLocationAbundanceNotesImages
Tessarolax [9] T. louellaeAn aporrhaid gastropod.

Crustaceans

GenusSpeciesLocationAbundanceNotesImages
Neocallichirus N. manningiA callianassid decapod.
Longusorbis L. cuniculosusA longusorbiid decapod.
Unnamed raninoid [13] UnnamedA raninoid decapod.

Paleoflora

Gymnosperms

GenusSpeciesLocationAbundanceNotesImages
Cunninghamia C. hornbyensisPermineralized wigs and leaves.A member of the extant Cupressaceae genus Cunninghamia. Closely resembles extant species. [14] Cunninghamia.jpg
Cycadeoidea [15] C. maccafferyiiA bennettitalean. Cycadeoidea fossil cropped.png

Angiosperms

GenusSpeciesLocationAbundanceNotesImages
Atli [16] A. moriniiStemA member of the Ranunculales with a liana-like growth habit.

See also

Footnotes

  1. 1 2 Martin-Silverstone, Elizabeth; Witton, Mark P.; Arbour, Victoria Megan; Currie, Philip J. (2016). "A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants". Royal Society Open Science. 3 (8): 160333. Bibcode:2016RSOS....360333M. doi: 10.1098/rsos.160333 . PMC   5108964 . PMID   27853614.
  2. 1 2 3 4 5 6 Fuchs; Beard; Tanabe; Ross, Dirk; G.; K.; R. (2007-08-17). "Coleoid cephalopods from the Late Cretaceous north eastern Pacific" (PDF). Seventh International Symposium 'Cephalopods - Present & Past'.: 131.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  3. Cappetta, Henri; Morrison, Kurt; Adnet, Sylvain (2021-08-03). "A shark fauna from the Campanian of Hornby Island, British Columbia, Canada: an insight into the diversity of Cretaceous deep-water assemblages" . Historical Biology. 33 (8): 1121–1182. Bibcode:2021HBio...33.1121C. doi:10.1080/08912963.2019.1681421. ISSN   0891-2963. S2CID   212878837.
  4. Cappetta, Henri; Morrison, Kurt; Adnet, Sylvain (2021-08-03). "A shark fauna from the Campanian of Hornby Island, British Columbia, Canada: an insight into the diversity of Cretaceous deep-water assemblages" . Historical Biology. 33 (8): 1121–1182. Bibcode:2021HBio...33.1121C. doi:10.1080/08912963.2019.1681421. ISSN   0891-2963. S2CID   212878837.
  5. Vullo, Romain; Buffetaut, Eric; Everhart, Michael J. (2012). "Reappraisal of Gwawinapterus beardi from the Late Cretaceous of Canada: a saurodontid fish, not a pterosaur" . Journal of Vertebrate Paleontology. 32 (5): 1198–1201. Bibcode:2012JVPal..32.1198V. doi:10.1080/02724634.2012.681078. ISSN   0272-4634. S2CID   129180570.
  6. McLachlan, Sandy M. S.; Kaiser, Gary W.; Longrich, Nicholas R. (2017-12-08). "Maaqwi cascadensis: A large, marine diving bird (Avialae: Ornithurae) from the Upper Cretaceous of British Columbia, Canada". PLOS ONE. 12 (12): e0189473. Bibcode:2017PLoSO..1289473M. doi: 10.1371/journal.pone.0189473 . ISSN   1932-6203. PMC   5722380 . PMID   29220405.
  7. Mayr, G. (2022). "Basic Terminology and the Broader Phylogenetic and Geological Framework)". Paleogene Fossil Birds. Springer Cham. pp. 3–27. doi:10.1007/978-3-030-87645-6_2.
  8. Dyke, Gareth; Wang, Xia; Kaiser, Gary (2011). Sues, Hans-Dieter (ed.). "Large fossil birds from a Late Cretaceous marine turbidite sequence on Hornby Island (British Columbia)" . Canadian Journal of Earth Sciences. 48 (11): 1489–1496. Bibcode:2011CaJES..48.1489D. doi:10.1139/e11-050. ISSN   0008-4077.
  9. 1 2 "PBDB". paleobiodb.org. Retrieved 2024-01-23.
  10. McLachlan, Sandy M. S.; Haggart, James W. (2018-12-08). "Reassessment of the late Campanian (Late Cretaceous) heteromorph ammonite fauna from Hornby Island, British Columbia, with implications for the taxonomy of the Diplomoceratidae and Nostoceratidae" . Journal of Systematic Palaeontology. 16 (15): 1247–1299. Bibcode:2018JSPal..16.1247M. doi:10.1080/14772019.2017.1381651. ISSN   1477-2019. S2CID   134663270.
  11. "PBDB". paleobiodb.org. Retrieved 2024-01-23.
  12. "PBDB". paleobiodb.org. Retrieved 2024-01-23.
  13. Nyborg, Torrey; Garassino, Alessandro; Vega, Francisco J.; Ross, Richard L. M. (2024-03-13). "A new fossil frog crab (Brachyura, Raninoidea) from the late Campanian of Hornby Island (British Columbia, Canada)" . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen: 209–217. doi:10.1127/njgpa/2023/1177.
  14. Brink, Kirstin S.; Stockey, Ruth A.; Beard, Graham; Wehr, Wesley C. (2009-05-01). "Cunninghamia hornbyensis sp. nov.: Permineralized twigs and leaves from the Upper Cretaceous of Hornby Island, British Columbia, Canada" . Review of Palaeobotany and Palynology. 155 (1): 89–98. Bibcode:2009RPaPa.155...89B. doi:10.1016/j.revpalbo.2009.03.005. ISSN   0034-6667.
  15. Rothwell, Gar W.; Stockey, Ruth A. (2002). "Anatomically Preserved Cycadeoidea (Cycadeoidaceae), with a Reevaluation of Systematic Characters for the Seed Cones of Bennettitales" . American Journal of Botany. 89 (9): 1447–1458. doi:10.3732/ajb.89.9.1447. ISSN   0002-9122. JSTOR   4124062. PMID   21665746.
  16. Smith, Selena Y.; Little, Stefan A.; Cooper, Ranessa L.; Burnham, Robyn J.; Stockey, Ruth A. (2013). "A Ranunculalean Liana Stem from the Cretaceous of British Columbia, Canada: Atli morinii gen. et sp. nov" . International Journal of Plant Sciences. 174 (5): 818–831. doi:10.1086/669925. ISSN   1058-5893. S2CID   84787422.

References