Northumberland Formation

Last updated
Northumberland Formation
Stratigraphic range: Late Cretaceous,
?Campanian–Maastrichtian
Type Geological formation
Unit of Nanaimo Group
Underlies Spray Formation
Location
Coordinates 49°36′N124°42′W / 49.6°N 124.7°W / 49.6; -124.7
Approximate paleocoordinates 56°06′N94°24′W / 56.1°N 94.4°W / 56.1; -94.4
CountryFlag of Canada (Pantone).svg  Canada

The Northumberland Formation is a Late Cretaceous (?Campanian-?Maastrichtian)-aged geologic formation in Canada. It belongs to the larger Nanaimo Group. Indeterminate bird and pterosaur fossils have been recovered from the formation, [1] as well as a potential gladius of Eromangateuthis . [2] An extensive diversity of shark teeth is known from the formation; many appear to be closely allied with modern deep-water shark taxa, suggesting a deep-water environment for the formation. [3] The most well-known exposures of the formation are on Hornby Island.

Contents

Vertebrate paleofauna

Cartilaginous fish [4]

Hybodontiformes

GenusSpeciesLocationAbundanceNotesImages
Meristodonoides M. sp.A hybodontid shark.

Hexanchiformes

GenusSpeciesLocationAbundanceNotesImages
Chlamydoselachus C. balliA chlamydoselachid shark, related to the modern frilled shark. Chlamydoselachus anguineus2.jpg
Rolfodon R. ludvigseniA chlamydoselachid shark.
R. cf. thompsoni
Dykeius D. garethiA very large chlamydoselachid shark.
Hexanchus H. microdonA cow shark related to the modern bluntnose sixgill shark. Hexanchus griseus santa rosa.jpg
Notidanodon N. pectinatusA cow shark. Hexanchidae species - Tooth.jpg
Xampylodon X. dentatusA large cow shark. Notidanodon.jpg
Protoheptranchias P. loweiA cow shark similar to the modern sharpnose sevengill shark.
Paraorthacodus P. rossiA paraorthacodontid shark. Paraorthacodus.jpg
Komoksodon K. kwutchakuthA komoksodontid shark.

Echinorhiniformes

GenusSpeciesLocationAbundanceNotesImages
Echinorhinus E. lapaoiA echinorhinid shark, related to the modern bramble shark. Echinorhinus brucus Mc Coy.jpg

Squaliformes

GenusSpeciesLocationAbundanceNotesImages
Eoetmopterus E. supercretaceusAn etmopterid shark similar to modern lantern sharks.
Squalus S. vondermarckiA dogfish shark, related to modern spurdogs. Squalus acanthias stellwagen.jpg
S. nicholsae
S. sp.
Centrosqualus C. mustardiA dogfish shark.
Protocentrophorus P. steviaeA dogfish shark.
Rhinoscymnus R. clarkiA sleeper shark, possibly included within Somniosus . Somniosus rostratus.png
Centroscymnus C. sp.A sleeper shark related to the modern Portuguese dogfish. Centroscymnus coelolepis norfanz.jpg
Squaliodalatias S. savoieiA dalatiid shark.
Hessinodon H. wardiA possibly dalatiid shark.

Pristiophoriformes

GenusSpeciesLocationAbundanceNotesImages
Pristiophorus P. smithiA sawshark. Pristiophorus japonicus cropped.jpg
P. pricei

Orectolobiformes

GenusSpeciesLocationAbundanceNotesImages
Plicatoscyllium P. cf. globidensA ginglymostomatid shark.
Hemiscyllium H. hermaniA bamboo shark. Hemiscyllium halmahera.jpg

Lamniformes

GenusSpeciesLocationAbundanceNotesImages
Carcharias C. domingueiA sand shark, related to the modern sand tiger shark. Grey Nurse Shark at Fish Rock Cave, NSW.jpg

Synechodontiformes

GenusSpeciesLocationAbundanceNotesImages
Synechodus S. derekiA palaeospinacid shark.

Carcharhiniformes

GenusSpeciesLocationAbundanceNotesImages
Scyliorhinidae indent.A catshark of uncertain affinities.
Florenceodon F. johnyiA florenceodontid shark.

Bony fish

GenusSpeciesLocationAbundanceNotesImages
Gwawinapterus G. beardiJaw.A saurodontid ichthyodectiform, initially identified as an istiodactylid pterosaur. [5]

Birds

GenusSpeciesLocationAbundanceNotesImages
Maaqwi [6] M. cascadensisCoracoid and wing bones.A large ornithuran, either a vegaviid or a procellariform. [7] Maaqwi.png
Enantiornithes indet. [8] An enanthiornithine avialan.

Pterosaurs

GenusSpeciesLocationAbundanceNotesImages
Azhdarchoidea indet. [1] A possibly azhdarchid pterosaur.

Squamates

GenusSpeciesLocationAbundanceNotesImages
Mosasauridae indet. [9] A mosasaur.

Invertebrate paleofauna

Molluscs

Cephalopods [10]

GenusSpeciesLocationAbundanceNotesImages
Baculites B. occidentalisA baculitid ammonite. Baculites fossil cropped.png
Fresvillia F. constrictaA baculitid ammonite.
Diplomoceras D. cylindraceumA diplomoceratid ammonite. Diplomoceras.jpg
D. cf. cylindraceum
Exiteloceras E. densicostatumA diplomoceratid ammonite. Exiteloceras jenneyi 1.jpg
E. bipunctatum
Phylloptychoceras P. horitaiA diplomoceratid ammonite.
Solenoceras S. exornatusA diplomoceratid ammonite.
S. cf. reesidei
Nostoceras N. adrotansA nostoceratid ammonite. Nostoceras species cropped.jpg
N. hornbyensis
N. aff. pauper
Enchoteuthis [2] E. sp.A muensterellid octopodiform. Enchoteuthis reconstruction.png
Eromangateuthis [2] E. soniae?A plesioteuthid octopodiform.
Cyrtobelus [11] C. hornbyenseA groenlandibelid spirulid. [2]
Actinosepia [2] A. canadensisAn actinosepiid vampyromorphid. [12]
Cirroteuthidae indet. [2] Gladius.A cirroteuthid octopus.

Gastropods

GenusSpeciesLocationAbundanceNotesImages
Tessarolax [9] T. louellaeAn aporrhaid gastropod.

Crustaceans

GenusSpeciesLocationAbundanceNotesImages
Neocallichirus N. manningiA callianassid decapod.
Longusorbis L. cuniculosusA longusorbiid decapod.
Unnamed raninoid [13] UnnamedA raninoid decapod.

Paleoflora

Gymnosperms

GenusSpeciesLocationAbundanceNotesImages
Cunninghamia C. hornbyensisPermineralized wigs and leaves.A member of the extant Cupressaceae genus Cunninghamia. Closely resembles extant species. [14] Cunninghamia.jpg
Cycadeoidea [15] C. maccafferyiiA bennettitalean. Cycadeoidea fossil cropped.png

Angiosperms

GenusSpeciesLocationAbundanceNotesImages
Atli [16] A. moriniiStemA member of the Ranunculales with a liana-like growth habit.

See also

Footnotes

  1. 1 2 Martin-Silverstone, Elizabeth; Witton, Mark P.; Arbour, Victoria Megan; Currie, Philip J. (2016). "A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants". Royal Society Open Science. 3 (8): 160333. Bibcode:2016RSOS....360333M. doi: 10.1098/rsos.160333 . PMC   5108964 . PMID   27853614.
  2. 1 2 3 4 5 6 Fuchs; Beard; Tanabe; Ross, Dirk; G.; K.; R. (2007-08-17). "Coleoid cephalopods from the Late Cretaceous north eastern Pacific" (PDF). Seventh International Symposium 'Cephalopods - Present & Past'.: 131.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  3. Cappetta, Henri; Morrison, Kurt; Adnet, Sylvain (2021-08-03). "A shark fauna from the Campanian of Hornby Island, British Columbia, Canada: an insight into the diversity of Cretaceous deep-water assemblages". Historical Biology. 33 (8): 1121–1182. Bibcode:2021HBio...33.1121C. doi:10.1080/08912963.2019.1681421. ISSN   0891-2963. S2CID   212878837.
  4. Cappetta, Henri; Morrison, Kurt; Adnet, Sylvain (2021-08-03). "A shark fauna from the Campanian of Hornby Island, British Columbia, Canada: an insight into the diversity of Cretaceous deep-water assemblages". Historical Biology. 33 (8): 1121–1182. Bibcode:2021HBio...33.1121C. doi:10.1080/08912963.2019.1681421. ISSN   0891-2963. S2CID   212878837.
  5. Vullo, Romain; Buffetaut, Eric; Everhart, Michael J. (2012). "Reappraisal of Gwawinapterus beardi from the Late Cretaceous of Canada: a saurodontid fish, not a pterosaur". Journal of Vertebrate Paleontology. 32 (5): 1198–1201. Bibcode:2012JVPal..32.1198V. doi:10.1080/02724634.2012.681078. ISSN   0272-4634. S2CID   129180570.
  6. McLachlan, Sandy M. S.; Kaiser, Gary W.; Longrich, Nicholas R. (2017-12-08). "Maaqwi cascadensis: A large, marine diving bird (Avialae: Ornithurae) from the Upper Cretaceous of British Columbia, Canada". PLOS ONE. 12 (12): e0189473. Bibcode:2017PLoSO..1289473M. doi: 10.1371/journal.pone.0189473 . ISSN   1932-6203. PMC   5722380 . PMID   29220405.
  7. Mayr, G. (2022). "Basic Terminology and the Broader Phylogenetic and Geological Framework)". Paleogene Fossil Birds. Springer Cham. pp. 3–27. doi:10.1007/978-3-030-87645-6_2.
  8. Dyke, Gareth; Wang, Xia; Kaiser, Gary (2011). Sues, Hans-Dieter (ed.). "Large fossil birds from a Late Cretaceous marine turbidite sequence on Hornby Island (British Columbia)". Canadian Journal of Earth Sciences. 48 (11): 1489–1496. Bibcode:2011CaJES..48.1489D. doi:10.1139/e11-050. ISSN   0008-4077.
  9. 1 2 "PBDB". paleobiodb.org. Retrieved 2024-01-23.
  10. McLachlan, Sandy M. S.; Haggart, James W. (2018-12-08). "Reassessment of the late Campanian (Late Cretaceous) heteromorph ammonite fauna from Hornby Island, British Columbia, with implications for the taxonomy of the Diplomoceratidae and Nostoceratidae". Journal of Systematic Palaeontology. 16 (15): 1247–1299. Bibcode:2018JSPal..16.1247M. doi:10.1080/14772019.2017.1381651. ISSN   1477-2019. S2CID   134663270.
  11. "PBDB". paleobiodb.org. Retrieved 2024-01-23.
  12. "PBDB". paleobiodb.org. Retrieved 2024-01-23.
  13. Nyborg, Torrey; Garassino, Alessandro; Vega, Francisco J.; Ross, Richard L. M. (2024-03-13). "A new fossil frog crab (Brachyura, Raninoidea) from the late Campanian of Hornby Island (British Columbia, Canada)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen: 209–217. doi:10.1127/njgpa/2023/1177.
  14. Brink, Kirstin S.; Stockey, Ruth A.; Beard, Graham; Wehr, Wesley C. (2009-05-01). "Cunninghamia hornbyensis sp. nov.: Permineralized twigs and leaves from the Upper Cretaceous of Hornby Island, British Columbia, Canada". Review of Palaeobotany and Palynology. 155 (1): 89–98. Bibcode:2009RPaPa.155...89B. doi:10.1016/j.revpalbo.2009.03.005. ISSN   0034-6667.
  15. Rothwell, Gar W.; Stockey, Ruth A. (2002). "Anatomically Preserved Cycadeoidea (Cycadeoidaceae), with a Reevaluation of Systematic Characters for the Seed Cones of Bennettitales". American Journal of Botany. 89 (9): 1447–1458. doi:10.3732/ajb.89.9.1447. ISSN   0002-9122. JSTOR   4124062. PMID   21665746.
  16. Smith, Selena Y.; Little, Stefan A.; Cooper, Ranessa L.; Burnham, Robyn J.; Stockey, Ruth A. (2013). "A Ranunculalean Liana Stem from the Cretaceous of British Columbia, Canada: Atli morinii gen. et sp. nov". International Journal of Plant Sciences. 174 (5): 818–831. doi:10.1086/669925. ISSN   1058-5893. S2CID   84787422.

Related Research Articles

<span class="mw-page-title-main">Carcharhiniformes</span> Order of sharks

Carcharhiniformes, commonly known as ground sharks, are the largest order of sharks, with over 270 species. They include a number of common types, such as catsharks, swellsharks, and requiem sharks.

<span class="mw-page-title-main">Hexanchiformes</span> Order of sharks

The Hexanchiformes are a primitive order of sharks, numbering just seven extant species in two families. Fossil sharks that were apparently very similar to modern sevengill species are known from Jurassic specimens.

<i>Cunninghamia</i> Genus of conifers

Cunninghamia is a genus of one or two living species of evergreen coniferous trees in the cypress family Cupressaceae. They are native to China, northern Vietnam and Laos, and perhaps also Cambodia. They may reach 50 m (160 ft) in height. In vernacular use, it is most often known as Cunninghamia, but is also sometimes called "China-fir". The genus name Cunninghamia honours Dr. James Cunningham, a British doctor who introduced this species into cultivation in 1702 and botanist Allan Cunningham.

<i>Chlamydoselachus</i> Genus of sharks

Chlamydoselachus is a genus of sharks and the sole extant member of the family Chlamydoselachidae, in the order Hexanchiformes. It contains two extant and four extinct species. The most widely known species still surviving is the frilled shark. It is known as a living fossil, along with Chlamydoselachus africana, also known as the southern African frilled shark, which is only found along coastal areas of South Africa. The only two extant species of this genus are deep-sea creatures which are typically weakened in areas closer to the surface. While the two extant species are similar in external appearance, they differ internally.

<span class="mw-page-title-main">Hornby Island</span> Island and human settlement in British Columbia, Canada

Hornby Island of British Columbia, Canada, is one of the two northernmost Gulf Islands, the other being Denman Island. It is located near Vancouver Island's Comox Valley,

<span class="mw-page-title-main">Istiodactylidae</span> Family of istiodactyliform pterosaurs

Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus Istiodactylus was discovered not to be a member of the genus Ornithodesmus.

The Toolebuc Formation is a geological formation that extends from Queensland across South Australia and the Northern Territory in Australia, whose strata date back to the Albian stage of the Early Cretaceous. Dinosaurs, pterosaurs, plesiosaurs, ichthyosaurs, protostegid turtles, sharks, chimaeroids and bony fish remains are among the fossils that have been recovered from the formation.

<i>Paraorthacodus</i> Extinct genus of sharks

Paraorthacodus is an extinct genus of shark. It a member of the family Paraorthacodontidae, which is either placed in Hexanchiformes or in Synechodontiformes. It is known from over a dozen named species spanning from the Early Jurassic to the Paleocene, or possibly Eocene. Almost all members of the genus are exclusively known from isolated teeth, with the exception of P. jurensis from the Late Jurassic-Early Cretaceous of Europe, which is known from full body fossils from the Late Jurassic of Germany, which suggest that juveniles had a robust body with a round head, while adults had large body sizes with a fusiform profile. There was only a single dorsal fin towards the back of the body without a fin spine. The dentition had teeth with a single large central cusp along with shorter lateral cusplets, which were designed for clutching. The teeth are distinguished from those of Synechodus by the lateral cusplets decreasing in size linearly away from the central cusp rather than exponentially as in Synechodus.

Gwawinapterus beardi is a species of saurodontid ray-finned fish from the Late Cretaceous period of British Columbia, Canada. While initially described as a very late-surviving member of the pterosaur family Istiodactylidae, further examination has cast doubt on the identification of the specimen as a pterosaur, and research published in 2012 identified the remains as having come from a saurodontid fish.

<i>Meristodonoides</i> Extinct genus of hybodont chondrichthyans

Meristodonoides is an extinct genus of hybodont known from the mid-late Cretaceous, with potential records dating back to the Jurassic. It is one of a number of hybodont genera composed of species formerly assigned to Hybodus.

The Tar Heel Formation, also known as the Coachman Formation in South Carolina, is a Late Cretaceous geologic formation in North Carolina and South Carolina, USA. It preserves fossils, including amber dating back to the Cretaceous period. A locality known as Phoebus Landing, has been dated to 78.5-77.1 Ma, and the formation has been overall dated to the early Campanian based on fossil pollen.

The Bladen Formation is a geologic formation from the Late Cretaceous of North Carolina and South Carolina, USA. It is known for a plethora of terrestrial and aquatic vertebrate fossils, including dinosaurs and mosasaurs. It appears to be roughly concurrent with the Tuscaloosa Formation of Alabama.

Notidanodon is an extinct genus of cow shark. Fossils ascribed to this genus are known from the Jurassic and Cretaceous periods. Recently, the genus underwent a major revision and was split into two after the erection of Xampylodon to accommodate the species X. dentatus, X. loozi, and X. brotzeni. The genus is now known only from New Zealand, Antarctica, Africa, and South America.

Adnetoscyllium is an extinct genus of bamboo shark from the Cretaceous period. It is currently monotypic, containing only the species A. angloparisensis. The genus is named for prominent paleoichthyologist, Dr. Sylvain Adnet. The specific epithet refers to the range which is thus far restricted to the Anglo-Paris Basin of France and the United Kingdom.

Hessinodon is an extinct genus of possible kitefin sharks that lived during the Late Cretaceous. It contains one valid species, H. wardi, which is known from four teeth from the Northumberland Formation of British Columbia. Its teeth are most similar to those of modern cookiecutter sharks.

Rolfodon is an extinct genus of shark in the family Chlamydoselachidae. It is closely related to the extant frilled sharks in the genus Chlamydoselachus, which it can be differentiated from by tooth morphology. It is named after late Canadian paleontologist Rolf Ludvigsen.

Dykeius is an extinct genus of large shark in the family Chlamydoselachidae. It contains a single known species, D. garethi, from the Late Cretaceous Northumberland Formation of Canada. The genus and species names honor paleontologist Gareth J. Dyke.

Komoksodon is an extinct genus of hexanchiform shark known from the Late Cretaceous to the Paleocene. It is the only member of the monotypic family Komoksodontidae.

<i>Xampylodon</i> Extinct genus of cow shark

Xampylodon is an extinct genus of cow shark. Fossils assigned to this genus are known from the Late Cretaceous and early Paleocene. Xampylodon was recently erected after a revision on the taxonomy of hexanchid fossil teeth, and includes four species, most of them previously included in Notidanodon.

Protocentrophorus is a genus of was a genus of dogfish shark that existed during the Cretaceous. Fossils have been found in the Tauragė County of Lithuania and British Columbia in Canada. Known from teeth, they were originally assigned to the genus Centrophorus.

References