Farlowichnus Temporal range: Late Cretaceous | |
---|---|
Trace fossil classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | Saurischia |
Clade: | Theropoda |
Ichnogenus: | † Farlowichnus Leonardi et al., 2023 |
Type ichnospecies | |
†Farlowichnus rapidus Leonardi et al., 2023 |
Farlowichnus is an ichnogenus of small theropod dinosaur footprint. It includes a single species, F. rapidus, known from prints found in the Early Cretaceous Botucatu Formation of Brazil. Farlowichnus is known from several fossil trackways that indicate that it was likely a cursorial animal that was well-adapted to desert environments. [1]
The Farlowichnus fossil material was discovered in sediments of the Botucatu Formation (São Bento Quarry) near Araraquara in São Paulo, Brazil. The holotype specimen, LPP-IC-0200, consists of a slab with four footprints forming a trackway. Two paratypes that preserve similar footprints were also assigned, consisting of LPP-IC-0231 and MCT-R-1954. [1]
In 2023, Farlowichnus rapidus was described by Giuseppe Leonardi, Marcelo A. Fernandes, Ismar S. Carvalho, Julia B. Schutzer, and Rafael C. da Silva as a new ichnogenus and ichnospecies of theropod footprints based on these fossil remains. The generic name, "Farlowichnus", combines a reference to James O. Farlow, a paleontologist who has studied ichnofossils, with the Greek "ίχνος" ("ichnos"), meaning "track". The specific name, "rapidus", references the hypothesized cursorial behavior and agile morphology of the taxon. [1]
Leonardi et al. (2023) described the general morphology of the Farlowichnus footprints as "waterdrop"-shaped. The impressions of the third digit of Farlowichnus are significantly longer than the second and fourth digits, which are proportionately much shorter. This morphology, as well as the observed long strides and high step angle, are associated with a cursorial behavior. Although the footprints are structurally tridactyl, or the imprints of three toes are visible, Farlowichnus was likely functionally monodactylous. The foot shape also indicates adaptations for running in soft, dry sand. [1]
The dinosaur trackmaker of the Farlowichnus footprints may have reached lengths of up to 3–3.5 metres (9.8–11.5 ft), as indicated by the larger size of one of the paratype specimens. The holotype trackmaker was smaller, at about 1.5 metres (4.9 ft) long, weighing around 12–15 kilograms (26–33 lb). [1]
While the type of theropod that produced the Farlowichnus is unknown, Leonardi et al. (2023) speculate that it may have been a relative of noasaurids. It may have had a body morphology comparable to the similarly aged Vespersaurus from Brazil's Rio Paraná Formation. The fossil material of Vespersaurus suggests that it was also monodactylous, a feature observed in the Farlowichnus trackways. [1] [2]
Farlowichnus was found in layers of the Botucatu Formation, which dates to the Berriasian–Hauterivian ages of the Early Cretaceous. Fossil remains of several indeterminate animals, including theropod and large ornithopod dinosaurs, mammals, lacertilian reptiles, and invertebrates, have also been described from the formation. The mammalian ichnotaxon Brasilichnium has also been identified from the formation. [3] [1]
Grallator ["GRA-luh-tor"] is an ichnogenus which covers a common type of small, three-toed print made by a variety of bipedal theropod dinosaurs. Grallator-type footprints have been found in formations dating from the Early Triassic through to the early Cretaceous periods. They are found in the United States, Canada, Europe, Australia, Brazil and China, but are most abundant on the east coast of North America, especially the Triassic and Early Jurassic formations of the northern part of the Newark Supergroup. The name Grallator translates into "stilt walker", although the actual length and form of the trackmaking legs varied by species, usually unidentified. The related term "Grallae" is an ancient name for the presumed group of long-legged wading birds, such as storks and herons. These footprints were given this name by their discoverer, Edward Hitchcock, in 1858.
Baurutitan is a genus of sauropod dinosaur that lived during the Late Cretaceous in what is now Brazil. The type species, Baurutitan britoi, was described in 2005 by Kellner and colleagues, although the fossil remains had already been discovered in 1957. Baurutitan is classified as a lithostrotian titanosaur, and is distinguished from related genera based on its distinctive caudal vertebrae. This South American dinosaur was found in the Serra da Galga Formation near Uberaba, in the Brazilian state of Minas Gerais.
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Tyrannosauripus is an ichnogenus of dinosaur footprint. It was discovered by geologist Charles "Chuck" Pillmore in 1983 and formally described by Martin Lockley and Adrian Hunt in 1994. This fossil footprint from northern New Mexico is 96 cm long and given its Late Cretaceous age, it very likely belonged to the giant theropod dinosaur Tyrannosaurus rex. In 2016 the size of this individual was estimated at 11.4 meters and 5.8-6.9 tonnes. Similar tridactyl dinosaur tracks in North America were discovered earlier and named Tyrannosauropus in 1971, but they were later recognized as hadrosaurid tracks and their description deemed inadequate, with Tyrannosauropus regarded as a nomen dubium. True footprints likely from Tyrannosaurus would not be found until the discovery of Tyrannosauripus. In 2007, a large tyrannosaurid track was found also in eastern Montana. In 2016, a probable fossil trackway of Tyrannosaurus was discovered in Wyoming.
Tyrannosauropus is a dubious ichnogenus of tridactyl dinosaur footprint from the Campanian of the Late Cretaceous of North America. Tyrannosauropus was named for a collection of footprints discovered on the ceiling of a cave in Utah which were suggested to have been made by Tyrannosaurus and informally labelled as "Tyrannosauripus" in 1924. These footprints would later be named by Haubold in 1971 as Tyrannosauropus petersoni, and attributed to Tyrannosaurus rex. However, Tyrannosauropus are Campanian in age, pre-dating the Maastrichtian age for Tyrannosaurus, and the morphology of the footprints more closely resembles those of hadrosaurid dinosaurs than those of theropods. Furthermore, in 1994 another footprint was described as likely belonging to Tyrannosaurus that matched it in both age and morphology and was named Tyrannosauripus. In the same publication, the description of Tyrannosauropus was deemed inadequate, with the holotype damaged and lost, and the ichnotaxon was declared undiagnostic and thus a nomen dubium.
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Hadrosaurichnoides is an Early Cretaceous ichnogenus of dinosaur footprint erected by María Lourdes Casanovas Cladellas. The original description attributed the ichnogenus to an ornithopod trackmaker, but in 2001 Lockley and Wright argued that they were actually left by theropods because the prints were longer and narrower than would be expected for ornithopod tracks.
The Botucatu Formation is an Aptian geologic formation of the Paraná and Pelotas Basins in southern Brazil and northern Uruguay. The formation is composed of quartzitic sandstones, deposited in an eolian environment. Fossil theropod tracks have been reported from the formation.
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