| Formiciinae | |
|---|---|
 | |
| Titanomyrma , with a rufous hummingbird for scale | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Hymenoptera |
| Family: | Formicidae |
| Subfamily: | † Formiciinae Lutz, 1986 |
| Type species | |
| Formicium berryi Westwood, 1854 | |
| Genera | |
| |
Formiciinae is an extinct subfamily of ants known from Eocene deposits in Europe and North America. [1]
The type genus is Formicium with the genus Titanomyrma being described in 2011. Formicium includes the described species which are known from fossil wings only. Formicium is known from three species. Formicium mirabile, named by Theodore D. A. Cockerell in 1920, and Formicium brodiei, named by John O. Westwood in 1854, are both known from fore wings found in the middle Eocene of Bournemouth, Dorset, England. [1] The third species named, Formicium berryi was named by Frank M. Carpenter in 1929 from the middle Eocene Claiborne Formation in Puryear, Tennessee, USA, though he misidentified the formation as the Wilcox Formation. F. berryi was the first described occurrence of the genus and, until 2011, the subfamily, in North America. [1] With the description of Titanomyrma, the two species already described from complete body specimens, Formicium giganteum and F. simillimum, were transferred to the new genus as Titanomyrma giganteum and T. simillimum respectively. Titanomyrma also contains a third species, T. lubei described in the same paper as the genus and which is the second member of the subfamily known from North America. [1]
While workers belonging to the subfamily have not been found, queens and males for T. giganteum and T. simillimum are known and T. lubei is known from a single queen. The average size for the queens and males in Titanomyrma is equal to that of some of the largest modern ants known. Only the queens of Dorylus wilverthi currently reach similar lengths, up to 52 millimetres (2.0 in) as the smallest species of Titanomyrma, T. lubei. [1]
Formiciinae members were restricted in habitat to living in regions which had a mesic wet climate and an average mean annual temperature of 20 °C (68 °F) or higher. This is similar to the restricted ranges of the largest species of modern ants. [1] The spread of the subfamily from Europe to North America is postulated to have been across the North Atlantic landbridges which were present in the Eocene. While the average temperatures for this route are thought to have been lower than the range needed for Formiciinae species, a series of warmer events throughout the Eocene are suggested as aides in the crossing. [1]
The family Gryllidae contains the subfamilies and genera which entomologists now term true crickets. Having long, whip-like antennae, they belong to the Orthopteran suborder Ensifera, which has been greatly reduced in the last 100 years : taxa such as the tree crickets, spider-crickets and their allies, sword-tail crickets, wood or ground crickets and scaly crickets have been moved or elevated to family level. The type genus is Gryllus and the first use of the family name "Gryllidae" was by Francis Walker.
Myrmeciinae is a subfamily of the Formicidae, ants once found worldwide but now restricted to Australia and New Caledonia. This subfamily is one of several ant subfamilies which possess gamergates, female worker ants which are able to mate and reproduce, thus sustaining the colony after the loss of the queen. The Myrmeciinae subfamily was formerly composed of only one genus, Myrmecia, but the subfamily was redescribed by Ward & Brady in 2003 to include two tribes and four genera. An additional three genera, one form genus, and 9 species were described in 2006 from the Early Eocene of Denmark, Canada, and Washington. Subsequently an additional fossil genus was moved from the family Rhopalosomatidae in 2018, and a new genus was described in 2021.
Formicium is an extinct collective genus of giant ants in the Formicidae subfamily Formiciinae. The genus currently contains three species, Formicium berryi, Formicium brodiei, and Formicium mirabile. All three species were described from Eocene aged sediments.
Prionomyrmex is an extinct genus of bulldog ants in the subfamily Myrmeciinae of the family Formicidae. It was first described by Gustav Mayr in 1868, after he collected a holotype worker of P. longiceps in Baltic amber. Three species are currently described, characterised by their long mandibles, slender bodies and large size. These ants are known from the Eocene and Late Oligocene, with fossil specimens only found around Europe. It is suggested that these ants preferred to live in jungles, with one species assumed to be an arboreal nesting species. These ants had a powerful stinger that was used to subdue prey. In 2000, it was suggested by Cesare Baroni Urbani that the living species Nothomyrmecia macrops and a species he described both belonged to Prionomyrmex, but this proposal has not been widely accepted by the entomological community. Instead, scientists still classify the two genera distinctive from each other, making Nothomyrmecia a valid genus.
Dinopanorpidae is a small family of extinct insects in the order Mecoptera (scorpionflies) that contains two genera and seven species.
Dinopanorpa is an extinct monotypic genus of scorpionfly that contains the single species Dinopanorpa megarche and is the type genus of the extinct family Dinopanorpidae. The genus is known from a single hindwing specimen, the holotype, currently deposited in the collections of the National Museum of Natural History, as number "69173", and which was first described by Dr Theodore D.A. Cockerell in 1924. The name is a combination of the Greek deino meaning "terrible" or "monstrous" and "Panorpa", the type genus of Panorpidae the family in which Dinopanorpa was first placed.
Titanomyrma is a genus of extinct giant ants which lived during the Eocene. The type species Titanomyrma gigantea and the smaller Titanomyrma simillima are known from the Eocene of Germany, while the third species Titanomyrma lubei, is known from Wyoming, United States. The presence of Titanomyrma in North America was considered to indicate "the first reported cross-Arctic dispersal by a thermophilic insect group". However a queen reported from Upland temperate shales in British Columbia raised questions on the exact thermophilic nature of the genus. The type species of this genus, T. gigantea, is the largest-known fossil or extant species of ant in the world.
Ypresiomyrma is an extinct genus of ants in the subfamily Myrmeciinae that was described in 2006. There are four species described; one species is from the Isle of Fur in Denmark, two are from the McAbee Fossil Beds in British Columbia, Canada, and the fourth from the Bol’shaya Svetlovodnaya fossil site in Russia. The queens of this genus are large, the mandibles are elongated and the eyes are well developed; a stinger is also present. The behaviour of these ants would have been similar to that of extant Myrmeciinae ants, such as solitary foraging for arthropod prey and never leaving pheromone trails. The alates were poor flyers due to their size, and birds and animals most likely preyed on these ants. Ypresiomyrma is not assigned to any tribe, and is instead generally regarded as incertae sedis within Myrmeciinae. However, some authors believe Ypresiomyrma should be assigned as incertae sedis within Formicidae.
Avitomyrmex is an extinct genus of bulldog ants in the subfamily Myrmeciinae which contains three described species. The genus was described in 2006 from Ypresian stage deposits of British Columbia, Canada. Almost all the specimens collected are queens, with an exception of a single fossilised worker. These ants are large, and the eyes are also large and well-developed; a sting is present in one species. The behaviour of these ants may have been similar to extant Myrmeciinae ants, such as foraging solitarily for arthropod prey and never leaving pheromone trails to food sources. Avitomyrmex has not been assigned to any tribe, instead generally being regarded as incertae sedis within Myrmeciinae. However, its identity as an ant has been challenged, although it is undoubtedly a hymenopteran insect.
Macabeemyrma is an extinct genus of bulldog ants in the subfamily Myrmeciinae containing the single species Macabeemyrma ovata, described in 2006 from Ypresian stage deposits of British Columbia, Canada. Only a single specimen is known; a holotype queen found preserved as a compression fossil. The specimen had no wings and small portions of its legs and eyes were faintly preserved. It was a large ant, reaching 25 millimetres (0.98 in) in length. This ants' behaviour would have been similar to that of extant Myrmeciinae ants, such as foraging singly in search for arthropod prey and nesting in soil or in trees. Macabeemyrma shows similarities to extinct ants in the genus Ypresiomyrma, and to the living Nothomyrmecia macrops, but has not been conclusively assigned to any tribe, instead generally regarded as incertae sedis within Myrmeciinae. However, the sole specimen lacks definitive traits, and its classification in Myrmeciinae, and even its identity as an ant, has been challenged.
Myrmeciites is an extinct form genus of bulldog ants in the subfamily Myrmeciinae of the family Formicidae, which contains three described species and two fossils not placed beyond the genus level. Described in 2006 from Ypresian stage deposits, all three of the described species and one unplaced fossil are from British Columbia, Canada, while the second unplaced fossil is from Washington State, USA. These ants were large, with the largest specimens collected reaching 3 centimetres (1.2 in). The behaviour of these ants would have been similar to extant Myrmeciinae ants, such as solitary foraging, nesting either in the soil or trees, and leaving no pheromone trail to food sources. Due to the poor preservation of these ants, their phylogenetic position among Myrmeciinae is unclear, and no type species has been designated. These ants are classified as incertae sedis in Myrmeciinae, but some writers have classified it as incertae sedis within the insect order Hymenoptera. This reclassification, however, has not been accepted; instead, Myrmeciites remains in Myrmeciinae.
The Allenby formation is a sedimentary rock formation in British Columbia which was deposited during the Ypresian stage of the Early Eocene. It consists of conglomerates, sandstones with interbedded shales and coal. The shales contain an abundance of insect, fish and plant fossils known from 1877 and onward, while the Princeton Chert was first indented in the 1950s and is known from anatomically preserved plants.
Prionomyrmecini is an ant tribe belonging to the subfamily Myrmeciinae established by William Morton Wheeler in 1915. Two members are a part of this tribe, the extant Nothomyrmecia and the extinct Prionomyrmex. The tribe was once considered a subfamily due to the similarities between Nothomyrmecia and Prionomyrmex, but such reclassification was not widely accepted by the scientific community. These ants can be identified by their long slender bodies, powerful stingers and elongated mandibles. Fossil Prionomyrmecini ants were once found throughout Europe, possibly nesting in trees and preferring jungle habitats. Today, Prionomyrmecini is only found in Australia, preferring old-growth mallee woodland surrounded by Eucalyptus trees. Nothomyrmecia workers feed on nectar and arthropods, using their compound eyes for prey and navigational purposes. Owing to their primitive nature, they do not recruit others to food sources or create pheromone trails. Nothomyrmecia colonies are small, consisting of 50 to 100 individuals.
Aphaenogaster mayri is an extinct species of ant in formicid subfamily Myrmicinae known from a series of Late Eocene fossils found in North America. A. mayri was one of two Aphaenogaster species described in a 1930 paper by Frank M. Carpenter.
Archimyrmex is an extinct genus of ant in the formicid subfamily Myrmeciinae, described by palaeoentomologist Theodore Cockerell in 1923. The genus contains four described species, Archimyrmex rostratus, Archimyrmex piatnitzkyi, Archimyrmex smekali and Archimyrmex wedmannae. Archimyrmex is known from a group of Middle Eocene fossils which were found in North America, South America, and Europe. The genus was initially placed in the subfamily Ponerinae, but it was later placed in Myrmeciinae; it is now believed to be the ancestor of the extant primitive genus Myrmecia from Australia. Despite this, Archimyrmex is not a member to any tribe and is regarded as incertae sedis within Myrmeciinae. However, some authors believe Archimyrmex should be assigned as incertae sedis within Formicidae. These ants can be characterised by their large mandibles and body length, ranging from 13.2 to 30 mm. They also have long, thin legs and an elongated mesosoma (thorax) and petiole.
Pachycondyla lutzi is an extinct species of ant in the formicid subfamily Ponerinae described by from fossils found in Europe. P. lutzi is one of six Lutetian Pachycondyla species.
Klondikia is an extinct hymenopteran genus in the ant family Formicidae with a single described species Klondikia whiteae. The species is solely known from the Early Eocene sediments exposed in northeast Washington state, United States. The genus is currently not placed into any ant subfamily, being treated as incertae sedis.
The paleofauna of the Eocene Okanagan Highlands consists of Early Eocene arthropods, vertebrates, plus rare nematodes and molluscs found in geological formations of the northwestern North American Eocene Okanagan Highlands. The highlands lake bed series' as a whole are considered one of the great Canadian Lagerstätten. The paleofauna represents that of a late Ypresian upland temperate ecosystem immediately after the Paleocene-Eocene thermal maximum, and before the increased cooling of the middle and late Eocene to Oligocene. The fossiliferous deposits of the region were noted as early as 1873, with small amounts of systematic work happening in the 1880-90s on British Columbian sites, and 1920-30s for Washington sites. Focus and more detailed descriptive work on the Okanagan Highlands site started in the last 1970's. Most of the highlands sites are preserved as compression-impression fossils in "shales", but also includes a rare permineralized biota and an amber biota.
Palaeorehniidae is an extinct family of katydid-like orthopterans that has been described from the fossil record. The family is known from the Paleocene to the end of the Eocene and has been described from North America and Scotland. Circumscription and placement of the group has changed several times since it was first described in 1939, with the group currently treated as a family that is incertae sedis in the suborder Ensifera. Five monotypic genera are assigned to the family.
Media related to Formiciinae at Wikimedia Commons
Data related to Formiciinae at Wikispecies