Furca Temporal range: Upper Ordovician | |
---|---|
Reconstruction of Furca bohemica | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | † Marrellomorpha |
Order: | † Marrellida |
Genus: | † Furca Fritsch, 1908 |
Type species | |
Furca bohemica Fritsch, 1908 | |
Species | |
Furca bohemicaFritsch, 1908 Contents |
Furca (Latin for "fork") is an extinct genus of marrellomorph arthropod known from the Sandbian stage (upper Ordovician period) of the Czech Republic, [1] with a single currently described species, Furca bohemica. A tentative additional species, "Furca mauretanica": was proposed for specimens discovered in Morocco, but this species remains a nomen nudum until formally published, and probably belongs in a new separate genus.
Furca is characterized by a broad head shield with three pairs of prominent spines: front (anterolateral), side (mediolateral) and rear (posterolateral). The head shield of Furca ranged from 21–31 mm (0.83–1.22 in) long and 18–29 mm (0.71–1.14 in) at maximum width. The mediolateral spines are long and strongly curved. The outline of the head shield possess a fringe of small, gently curving secondary spines, between 1.5–7 mm (0.059–0.276 in) long. Appendages are unknown from fossils. [2]
Furca fossils have been found in sediments indicative of shallow marine habitats. Since appendages and other body parts are unknown, no firm conclusions can be made of the biology of Furca. However, comparisons to other marrellomorphs and living arthropods such as horseshoe crabs suggest a benthic marine lifestyle: dwelling on the sea floor. [2]
With its unusual anatomy, Furca has a colorful taxonomic history. The first specimens were discovered by Joachim Barrande, who named them Furca bohemica (Latin for "Bohemian fork") but did not formally publish descriptions. The specimens were not illustrated until 1847, when they were interpreted as the hypostome of the trilobite Prionocheilus . Antonin Fritsch was the first to formally describe Furca as a distinct taxon, interpreting F. bohemica as a larval echinoderm in 1908. [3] [4] In 1919, F. bohemica was, for the first time, interpreted as an arthropod. A second species, F. pilosa was described in 1999, as well as an unnamed species simply called "Furca sp.". In 2013, Rak and colleagues suggested that F. pilosa and all previously named species were synonyms of Furca bohemica. [2]
A tentative additional species, "Furca mauretanica" from the Floian stage Fezouata Formation of Morocco, was proposed in a doctoral thesis in 2006 [5] and subsequently referred to as "probably belonging to the genus Furca". [6] However, the name has yet to be formally published and so remains a nomen nudum . Phylogenetic analyses have recovered it as distinct marrelid taxon not closely related to Furca bohemica, and thus probably should be placed in a separate genus. [7]
Furca bohemica is assigned to the marrellomorph clade Marrellida. [2] While historicaly placed in Mimetasteridae alongside Mimetaster , other studies have disputed this placement. [7]
Marrella is an extinct genus of marrellomorph arthropod known from the Middle Cambrian of North America and Asia. It is the most common animal represented in the Burgess Shale of British Columbia, Canada, with tens of thousands of specimens collected. Much rarer remains are also known from deposits in China.
Nektaspida is an extinct order of non-mineralised artiopodan arthropods. They are known from the mid-Cambrian to the upper Silurian. Originally classified as trilobites, which they superficially resemble, they are now placed as close relatives as members of the Trilobitomorpha within Artiopoda. The order is divided into three major families; Emucarididae, Liwiidae, and Naraoiidae.
Skania is a Cambrian fossil arthropod. The type species, S. fragillis, is known from the Burgess Shale of British Columbia, Canada. A second possible species "S." sundbergi is known from the Kaili Formation of China, but its placement within the genus has been questioned.
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.
Anomalocaris is an extinct genus of radiodont, an order of early-diverging stem-group arthropods.
Megalograptus is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Megalograptus have been recovered in deposits of Katian age in North America. The genus contains five species: M. alveolatus, M. ohioensis, M. shideleri, M. welchi and M. williamsae, all based on fossil material found in the United States. Fossils unassigned to any particular species have also been found in Canada. The generic name translates to "great writing" and originates from the mistaken original belief that Megalograptus was a type of graptolite, often given names ending with -graptus.
Cheloniellida is a taxon of extinct Paleozoic arthropods. As of 2018, 7 monotypic genera of cheloniellids had been formally described, whose fossils are found in marine strata ranging from Ordovician to Devonian in age. Cheloniellida has a controversial phylogenetic position, with previous studies associated it as either a member or relative of various fossil and extant arthropod taxa. It was later accepted as a member of Vicissicaudata within Artiopoda.
Flexicalymene Shirley, 1936. is a genus of trilobites belonging to the order Phacopida, suborder Calymenina and Family Calymenidae. Flexicalymene specimens can be mistaken for Calymene, Gravicalymene, Diacalymene and a few other Calymenina genera. They are used as an index fossil in the Ordovician. Ohio and North America are particularly known for being rich with Flexicalymene fossils.
Choia is a genus of extinct demosponge ranging from the Cambrian until the Lower Ordovician periods. Fossils of Choia have been found in the Burgess Shale in British Columbia; the Maotianshan shales of China; the Wheeler Shale in Utah; and the Lower Ordovician Fezouata formation. It was first described in 1920 by Charles Doolittle Walcott.
Chasmataspidids, sometime referred to as chasmataspids, are a group of extinct chelicerate arthropods that form the order Chasmataspidida. Chasmataspidids are probably related to horseshoe crabs (Xiphosura) and/or sea scorpions (Eurypterida), with more recent studies suggest that they form a clade (Dekatriata) with Eurypterida and Arachnida. Chasmataspidids are known sporadically in the fossil record through to the mid-Devonian, with possible evidence suggesting that they were also present during the late Cambrian. Chasmataspidids are most easily recognised by having an opisthosoma divided into a wide forepart (preabdomen) and a narrow hind part (postabdomen) each comprising 4 and 9 segments respectively. There is some debate about whether they form a natural group.
Areia is a genus of cheirurid phacopid trilobite which existed during the Ordovician in what is now the Czech Republic. It was described by Barrande in 1872, and the type and only species is Areia bohemica.
Marrellomorpha are an extinct group of arthropods known from the Cambrian to the Early Devonian. They lacked mineralised hard parts, so are only known from areas of exceptional preservation, limiting their fossil distribution. The best known member is Marrella, with thousands of specimens found in the Cambrian aged Burgess Shale of Canada. The group is divided up into two major orders, Marrellida and Acercostraca. Marrellida is recognised by the possession of head shields with two or three pairs of elongate spine-like projections, and three pairs of uniramous appendages on the cephalon, while Acercostraca generally have large ovoid carapaces that cover the entire upper half of the body, and five pairs of uniramous cephalic appendages. Both groups have unbranched antennules and a segmented trunk with biramous appendages. Recent research has suggested the previously enigmatic Cambrian trilobite-like arthropods Skania and Primicaris belong to this group. Their phylogenetic position is uncertain, various studies have alternatively placed them in the Arachnomorpha as relatives of Artiopoda, as related to Mandibulata, or as stem group euarthropods.
Mimetaster is an extinct genus of marrellomorph arthropod. The type species, Mimetaster hexagonalis is known from the Lower Devonian (Pragian-Emsian) Hunsrück Slate, and amongst the most common arthropods from the locality, with over 120 specimens including three juveniles.
Cyclopyge is a genus of small to average size trilobites that lived during the Ordovician. Like all members of the family Cyclopigidae, it has very large convex eyes, that cover most of the free cheeks, and in some species touch each other. The eyes almost touch the large glabella. The occipital ring has merged with the rest of the glabella. The glabella does not extend into a frontal thorn. The cephalon lacks genal spines. The 6 thorax segments have short pleurae. The pygidium is rather large, and often rather effaced. These are features that also occur in other Cyclopygidae, and are indications of a pelagic lifestyle.
Aegirocassis is an extinct genus of giant radiodont arthropod belonging to the family Hurdiidae that lived 480 million years ago during the early Ordovician in the Fezouata Formation of Morocco. It is known by a single species, Aegirocassis benmoulai. Van Roy initiated scientific study of the fossil, the earliest known of a "giant" filter-feeder discovered to date. Aegirocassis is considered to have evolved from early predatory radiodonts. This animal is characterized by its long, forward facing head sclerite, and the endites on its frontal appendages that bore copious amounts of baleen-like auxiliary spines. This animal evolving filter-feeding traits was most likely a result of the Great Ordovician Biodiversification Event, when environmental changes caused a diversification of plankton, which in turn allowed for the evolution of new suspension feeding lifeforms. Alongside the closely related Pseudoangustidontus, an unnamed hurdiid from Wales, the middle Ordovician dinocaridid Mieridduryn, and the Devonian hurdiid Schinderhannes this radiodont is one of the few dinocaridids known from post-Cambrian rocks.
Strabopidae is the only family of the order Strabopida, an extinct group of arthropods known from the Cambrian period.
Cryptolithus is a genus of extinct epifaunal, suspension-feeding, trinucleid trilobites that lived during the Ordovician period. They were mostly blind. They are found in the United States, Canada, Venezuela, the United Kingdom, France, the Czech Republic, Morocco and Turkey.
The Letná Formation is a Late Ordovician geologic formation of the Prague Basin, Bohemian Massif in the Czech Republic. The formation crops out in the Czech capital, more specifically at Letná Hill, after which the formation is named. The type locality is located at Malá Strana, Holešovice district.
Tomlinsonus is an extinct genus of marellomorph arthropod known from the Late Ordovician (Katian) Kirkfield Formation of Ontario, Canada. It is a member of Marellida, and closely related to Mimetaster.