Gymnocarpium robertianum | |
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Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Division: | Polypodiophyta |
Class: | Polypodiopsida |
Order: | Polypodiales |
Suborder: | Aspleniineae |
Family: | Cystopteridaceae |
Genus: | Gymnocarpium |
Species: | G. robertianum |
Binomial name | |
Gymnocarpium robertianum | |
Synonyms | |
Dryopteris robertiana(Hoffm.) C. Chr. Contents |
Gymnocarpium robertianum, the limestone fern [1] or scented oakfern, [2] is a fern of the family Cystopteridaceae.
Gymnocarpium robertianum has small (10–50 cm), deltate, two- to three-pinnate fronds. Fronds arise from creeping rhizomes and have long, delicate rachis. The sori are borne in round clumps on the underside of the blade and lack an indusium. This species differs from the closely related G. dryopteris in having a densely glandular rachis as well as a more sparsely glandular underside to the blade. [3]
Gymnocarpium robertianum is thought to hybridise with G. appalachianum giving rise to Gymnocarpium × heterosporum W. H. Wagner . This hybrid was only known from Pennsylvania where it has now been eradicated. The hybrid between G. robertianum and G. dryopteris is called Gymnocarpium × achriosporumSarvela. This taxon is known from Sweden and Quebec. [4]
Gymnocarpium robertianum is a circumboreal species with populations in Europe, North America and the Caucasus Mountains. [4]
This species is protected in Illinois and Michigan [5] and is protected under the Flora Protection Order in Ireland. [6]
This species is a calcicole and as such is restricted to alkaline rich areas. In the British Isles its preferential habitat is grykes in limestone pavement. In Michigan the species is most frequent in Thuja occidentalis swamps. [7]
Asplenium platyneuron, commonly known as ebony spleenwort or brownstem spleenwort, is a fern native to North America east of the Rocky Mountains. It takes its common name from its dark, reddish-brown, glossy stipe and rachis, which support a once-divided, pinnate leaf. The fertile fronds, which die off in the winter, are darker green and stand upright, while the sterile fronds are evergreen and lie flat on the ground. An auricle at the base of each pinna points towards the tip of the frond. The dimorphic fronds and alternate, rather than opposite, pinnae distinguish it from the similar black-stemmed spleenwort.
Adiantum capillus-veneris, the Southern maidenhair fern, black maidenhair fern, maidenhair fern, and venus hair fern, is a species of ferns in the genus Adiantum and the family Pteridaceae with a subcosmopolitan worldwide distribution. It is cultivated as a popular garden fern and houseplant.
Dryopteris expansa, the alpine buckler fern, northern buckler-fern or spreading wood fern, is a species of perennial fern native to cool temperate and subarctic regions of the Northern Hemisphere, south at high altitudes in mountains to Spain and Greece in southern Europe, to Japan in eastern Asia, and to central California in North America. It prefers cool, moist mixed or evergreen forests and rock crevices on alpine slopes, often growing on rotting logs and tree stumps and rocky slopes. It is characteristically riparian in nature, and is especially associated with stream banks.
Dryopteris filix-mas, the male fern, is a common fern of the temperate Northern Hemisphere, native to much of Europe, Asia, and North America. It favours damp shaded areas in the understory of woodlands, but also shady places on hedge-banks, and rocks, and screes. Near the northern limit of its distribution it prefers sunny, well-drained sites. It is much less abundant in North America than in Europe.
Gymnocarpium dryopteris, the western oakfern, common oak fern, oak fern, or northern oak fern, is a deciduous fern of the family Cystopteridaceae. It is widespread across much of North America and Eurasia. It has been found in Canada, the United States, Greenland, China, Japan, Korea, Russia, and most of Europe. It is a seedless, vascular plant that reproduces via spores and have a life cycle with alternating, free-living sporophyte and gametophyte phases.
Polystichum aculeatum, the hard shield-fern, is an evergreen fern native to Europe. It is most abundant in upland regions of the British Isles and western France, where it benefits from the combination of mild winters and moist summers, but also occurs more locally across most of Europe except northern Scandinavia, northern Russia; in the Mediterranean region it is confined to high altitudes. It grows on steep slopes in woodlands. it is sometimes considered an indicator of the presence of ancient woodlands.
Dryopteris affinis, the scaly male fern or golden-scaled male fern, is a fern native to western and southern Europe and southwestern Asia. It is most abundant on moist soils in woodlands in areas with high humidity, such as the British Isles and western France. In the Mediterranean region and the Caucasus it is confined to high altitudes.
Dryopteris marginalis, vernacularly known as the marginal shield fern or marginal wood fern, is a perennial species of fern found in damp shady areas throughout eastern North America, from Texas to Minnesota and Newfoundland. It favors moderately acid to circumneutral soils in cooler areas but is fairly drought-resistant once established. In the warmer parts of its range, it is most likely to be found on north-facing non-calcareous rock faces. It is common in many altitudes throughout its range, from high ledges to rocky slopes and stream banks. Marginal wood fern's name derives from the fact that the sori are located on the margins, or edges of the leaflets.
Dryopteris carthusiana is a perennial species of fern native to damp forests throughout the Holarctic Kingdom. It is known as the narrow buckler-fern in the United Kingdom, and as the spinulose woodfern in North America.
Dryopteris cristata is a perennial species of fern native to wetlands throughout the Northern Hemisphere. It is known as crested wood fern, crested buckler-fern or crested shieldfern. This plant is a tetraploid species of hybrid origin, one parent being Dryopteris ludoviciana and the other being the unknown, apparently extinct species, dubbed Dryopteris semicristata, which is also one of the presumed parents of Dryopteris carthusiana. D. cristata in turn is one of the parents of Dryopteris clintoniana, another fern of hybrid origin.
Lemna trisulca L. is a species of aquatic plants in the arum family Araceae. It has a subcosmopolitan distribution. Unlike other duckweeds, it has submerged rather than floating fronds, except when flowering or fruiting. Also unlike other duckweeds, a large number of fronds remain attached to each other at a time.
Asplenium trichomanes, the maidenhair spleenwort, is a small fern in the spleenwort genus Asplenium. It is a widespread and common species, occurring almost worldwide in a variety of rocky habitats. It is a variable fern with several subspecies.
Botrychium lunaria is a species of fern in the family Ophioglossaceae known by the common name moonwort or common moonwort. It is the most widely distributed moonwort, growing throughout the Northern Hemisphere across Eurasia and from Alaska to Greenland, as well as temperate parts of the Southern Hemisphere.
Dryopteris intermedia, the intermediate wood fern or evergreen wood fern, is a perennial, evergreen wood fern native to eastern North America. It is a diploid species, and is the parent of several species of hybrid origin, including Dryopteris carthusiana. Other common names for this species include intermediate shield fern, fancy wood fern, fancy fern, glandular wood fern, American shield fern and common wood fern.
Pellaea atropurpurea, commonly known as purple-stem cliffbrake or just purple cliffbrake, is a fern native to North and Central America. Brake is an old word for fern, related to the word bracken. Like many other members of the Pteridaceae, it is a rock plant, needing a calcareous substrate.
Gymnocarpium is a small genus of ferns, called oak ferns. It was once placed with various other groups, including the dryopteroid ferns and the athyrioid ferns. Cladistic analysis has demonstrated that Gymnocarpium and Cystopteris form a natural but relatively primitive clade that is basal to the asplenioid, thelypterioid, and athyrioid ferns comprehensively.
Polystichum lonchitis is a species of fern known by the common name northern hollyfern, or simply holly-fern. It is native to much of the Northern Hemisphere from Eurasia to Alaska to Greenland and south into mountainous central North America. It has stiff, glossy green, erect fronds and grows in moist, shady, rocky mountain habitats.
Dryopteris dilatata, the broad buckler-fern, is a robust species of deciduous or semievergreen fern in the family Dryopteridaceae, native to Europe, particularly western and central Europe. In southern Europe, it is mostly found in mountainous regions. It is also found between the Black Sea and the Caspian Sea. It grows to 90 cm (35 in) tall by 120 cm (47 in) wide, with dark green tripinnate fronds, the ribs covered in brown scales.
Hymenophyllum tunbrigense, the Tunbridge filmy fern or Tunbridge filmy-fern, is a small, fragile perennial leptosporangiate fern which forms large dense colonies of overlapping leaves from creeping rhizomes. The common name derives from the leaves which are very thin, only a single cell thick, and translucent, giving the appearance of a wet film. The evergreen fronds are bipinnatifid, deeply and irregularly dissected, about 3 to 6 cm long, 2 cm across with dark winged stipes. In contrast to the similar H. wilsonii the fronds are more divided, flattened, appressed to the substrate and tend to have a bluish tint.
Dryopteris aemula, the hay-scented buckler-fern or hay-scented fern, is a species of perennial leptosporangiate fern.