Gymnocarpium

Gymnocarpium
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Order:	Polypodiales
Suborder:	Aspleniineae
Family:	Cystopteridaceae
Genus:	Gymnocarpium Newm. 1851
Type species
Gymnocarpium dryopteris (von Linné) Newman
Species
See text
Synonyms
Carpogymnia(Fuchs ex Janch.) Löve & Löve ×CystocarpiumFraser-Jenkins CurraniaCopeland

Gymnocarpium is a small genus of ferns, called oak ferns. It was once placed with various other groups, including the dryopteroid ferns and the athyrioid ferns. Cladistic analysis has demonstrated that Gymnocarpium and Cystopteris form a natural but relatively primitive clade that is basal to the asplenioid, thelypterioid, and athyrioid ferns comprehensively.

Gymnocarpium sori are small, round and naked. These ferns generally have a slender, creeping rhizome under the surface of the ground, and fairly thin-textured fronds.

Phylogeny

This genus includes the following species: ^[1]

Phylogeny from Fern Tree of Life [2] [3]

Other species include:

Gymnocarpium G. robertianum (Hoffm.) Newman (Limestone/scented oak fern) ×Cystocarpium × roskamianumFraser-Jenkins G. ×brittonianum(Sarvela) Pryer & Haufler G. disjunctum (Ruprecht) Ching (Pacific oak fern) G. dryopteris (von Linné) Newman (Western/northern oak fern) G. oyamense (Baker) Ching G. jessoense(Koidzumi) Koidzumi G. remotepinnatum(Hayata) Ching G. appalachianumPryer & Haufler (Appalachian oak fern) G. continentale(Petrov) Pojark. (Nahanni oak fern)

G. ×achriosporumSarvela G. ×bipinnatifidumMiyam. G. fedtschenkoanum G. ×intermediumSarvela

References

1. "Gymnocarpium dryopteris (L.)". Plants of the World Online. Royal Botanic Gardens, Kew. Retrieved 7 August 2025.
2. Nitta, Joel H.; Schuettpelz, Eric; Ramírez-Barahona, Santiago; Iwasaki, Wataru; et al. (2022). "An Open and Continuously Updated Fern Tree of Life". Frontiers in Plant Science. 13: 909768. doi: 10.3389/fpls.2022.909768 . PMC   9449725 . PMID   36092417.
3. "Tree viewer: interactive visualization of FTOL". FTOL v1.7.0 (GenBank release 261). 2024. Retrieved 1 May 2025.

External links

• Gymnocarpium world species list Archived 2018-02-02 at the Wayback Machine