Homolidae | |
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Paromola cuvieri | |
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Family: | Homolidae De Haan, 1839 |
The family Homolidae, known as carrier crabs [1] or porter crabs, [2] contains 14 genera of marine crabs. They mostly live on the continental slope and continental shelf, and are rarely encountered. [3] Members of the Homolidae have their fifth pereiopods (last pair of walking legs) in a sub-dorsal position, which allows them to hold objects in place over the rear half of the carapace. [3] The objects carried include sponges, black corals and gorgonians, and this behaviour may be a defence mechanism against predators. [3] Some species have been observed carrying living sea urchins in a symbiotic relationship which allows them to benefit from the protection of the urchin's dangerous spikes. [4] [5] [6] [7]
A total of 14 genera are currently recognised in the family: [8]
The Decapoda or decapods are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crabs, lobsters, crayfish, shrimp and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp and Anomura including hermit crabs, porcelain crabs, squat lobsters making up the bulk of the remainder. The earliest fossil decapod is the Devonian Palaeopalaemon.
Xanthidae is a family of crabs known as gorilla crabs, mud crabs, pebble crabs or rubble crabs. Xanthid crabs are often brightly coloured and are highly poisonous, containing toxins which are not destroyed by cooking and for which no antidote is known. The toxins are similar to the tetrodotoxin and saxitoxin produced by puffer fish, and may be produced by bacteria in the genus Vibrio living in symbiosis with the crabs, mostly V. alginolyticus and V. parahaemolyticus.
Portunidae is a family of crabs which contains the swimming crabs.
Xanthoidea is a superfamily of crabs, comprising the three families Xanthidae, Panopeidae and Pseudorhombilidae. Formerly, a number of other families were included in Xanthoidea, but many of these have since been removed to other superfamilies. These include Carpilioidea, Eriphioidea, Hexapodoidea, Pilumnoidea and Trapezioidea. Even in this reduced state, Xanthoidea remains one of the most species-rich superfamilies of crabs.
The Paguridae are a family of hermit crabs of the order Decapoda. This family contains 542 species in over 70 genera:
Lybia is a genus of small crabs in the family Xanthidae. Their common names include boxer crabs, boxing crabs and pom-pom crabs. They are notable for their mutualism with sea anemones, which they hold in their claws for defense. In return, the anemones get carried around which may enable them to capture more food particles with their tentacles. Boxer crabs use at least three species of anemones, including Bundeopsis spp. and Triactis producta. The bonding with the anemone is not needed for survival, however, and boxer crabs have frequently been known to live without them, sometimes substituting other organisms such as sponges and corals for the sea anemones.
Homolodromiidae is a family of crabs, the only family in the superfamily Homolodromioidea. In contrast to other crabs, including the closely related Homolidae, there is no strong linea homolica along which the exoskeleton breaks open during ecdysis. The family comprises two genera, Dicranodromia, which has 18 species, and Homolodromia, with five species.
Hexapodidae is a family of crabs, the only family in the superfamily Hexapodoidea. It has traditionally been treated as a subfamily of the family Goneplacidae, and was originally described as a subfamily of Pinnotheridae. Its members can be distinguished from all other true crabs by the reduction of the thorax, such that only seven sternites are exposed, and only four pairs of pereiopods are present. Not counting the enlarged pair of claws, this leaves only six walking legs, from which the type genus Hexapus, and therefore the whole family, takes its name. Some anomuran "crabs", such as porcelain crabs and king crabs also have only four visible pairs of legs. With the exception of Stevea williamsi, from Mexico, all the extant members are found either in the Indo-Pacific oceans, or around the coast of Africa.
Percnon gibbesi is a species of crab. It is one of at least two species commonly called "Sally Lightfoot", and is also referred to as the nimble spray crab or urchin crab. It has been described as "the most invasive decapod species to enter the Mediterranean".
Pseudozioidea is a superfamily of crabs, formerly treated in the Eriphioidea, Carpilioidea, Xanthoidea, Pilumnoidea and Goneplacoidea. A number of fossils from the Eocene onwards are known from the family Pseudoziidae. Eleven genera are recognised in three families:
Homola barbata is a species of crab in the family Homolidae.
Drachiella is a genus of crabs in the family Aethridae, containing one fossil species, and the following species:
Xantho is a genus of crabs in the family Xanthidae, containing five extant species, all restricted to the north-east Atlantic Ocean and Mediterranean Sea, although Xantho granulicarpis is not universally recognised as a separate species from Xantho hydrophilus:
Leucosiidae is a family of crabs containing three subfamilies and a number of genera incertae sedis:
Demania is a genus of crabs in the family Xanthidae, containing the following species:
Around 65 species of crab occur in the waters of the British Isles. All are marine, with the exception of the introduced Chinese mitten crab, Eriocheir sinensis, which occurs in fresh and brackish water. They range in size from the deep-water species Paromola cuvieri, which can reach a claw span of 1.2 metres, to the pea crab, which is only 4 mm (0.16 in) wide and lives inside mussel shells.
Danièle Guinot is a French biologist, an emeritus professor at the Muséum national d'histoire naturelle in France, known for her research on crabs.
Rochinia is a genus of crab in the family Epialtidae, containing the following species:
Mathildellidae is family of crabs belonging to the superfamily Goneplacoidea, containing the following genera: