Human hair color is the pigmentation of human hair follicles and shafts due to two types of melanin: eumelanin and pheomelanin. Generally, the more melanin present, the darker the hair. Its tone depends on the ratio of black or brown eumelanin to yellow or red pheomelanin. Melanin levels can vary over time, causing a person's hair color to change, and one person can have hair follicles of more than one color. Some hair colors are associated with some ethnic groups because of the observed higher frequency of particular hair colors within their geographical region, e.g. straight, dark hair amongst East Asians, Southeast Asians, Polynesians, some Central Asians, and Native Americans; a large variety of dark, fair, curly, straight, wavy or bushy amongst Europeans, West Asians, some Central Asians, and North Africans; and curly, dark, and uniquely helical hair amongst Sub Saharan Africans. Bright red hair is found in some European populations, and hair turns gray, white, or "silver" with age. [ citation needed ]
The full genetic basis of hair color is complex and not fully understood. [1] Regulatory DNA is believed to be closely involved in pigmentation in humans in general, [2] and a 2011 study by Branicki et al. identified 13 DNA variations across 11 different genes that could be used to predict hair color. [3]
Two types of pigment give hair its color, black-brown eumelanin and reddish-brown/reddish-yellow [4] pheomelanin, synthesized by melanocytes. [5] Inside the melanocytes, tyrosine is converted into L-DOPA and then L-dopaquinone, which in turn is formed into pheomelanin or eumelanin. [6]
Different hair color phenotypes arise primarily as a result of varying ratios of these two pigments in the human population, [5] although Europeans show the greatest range in pigmentation overall. [7] In addition, other genetic and environmental factors can affect hair color in humans; for instance, mutations in the melanocortin 1 receptor (MC1R) gene can lead to red or auburn hair, [5] and exposure to ultraviolet radiation can damage hair and alter its pigmentation. [8] Ultraviolet radiation (UV radiation) triggers greater synthesis of several compounds, including pro-opiomelanocortin (POMC), α-MSH, and ACTH, the result being increased eumelanin production. [6] UV radiation most commonly comes from the sun, and thus populations from places closer to the equator tend to have darker hair, [6] because eumelanin is generally more photoprotective than pheomelanin. [4]
Pheomelanin colors hair orange and red. Eumelanin, which has two subtypes of black or brown, determines the darkness of the hair color; [4] more black eumelanin leads to blacker hair and more brown eumelanin to browner hair. [6] All human hair has some amount of both pigments. [9] Over 95% of melanin content in black and brown hair is eumelanin. [9] Pheomelanin is generally found in elevated concentrations in blond and red hair, [4] representing about one-third of total melanin content. [9] If there is no black eumelanin, the result is strawberry blond. [6] Blond hair results from small amounts of brown eumelanin with no black eumelanin. [6]
Blond | Dark blond | Medium brown | Dark brown | Black | Reddish-brown | Red | Gray | White |
Natural hair color can be black, brown, blonde and red. [10]
The Fischer–Saller scale, named after Eugen Fischer and Karl Saller is used in physical anthropology and medicine to determine the shades of hair color. The scale uses the following designations: A (very light blond), B to E (light blond), F to L (blond), M to O (dark blond), P to T (light brown to brown), U to Y (dark brown to black) and Roman numerals I to IV (red) and V to VI (red-blond). [11]
Black hair or jet black hair is the darkest hair color. It has large amounts of eumelanin and is denser than other hair colors and is the commonly seen hair color in Asia and Africa due the fact that the people in these regions tend to have lower levels of tyrosinase in their bodies. Black eumelanin secretion causes the hair to turn black, which indicates that the MC1R is in the active state. Jet black hair, the darkest shade will not have a warm, neutral tone but a sheen which can seem almost blue, like the iridescence of a raven's wing; hence, sometimes referred to as raven-black. Jet black hair appears to have reflective silver color in bright sunlight. [12] [13]
Brown hair is the second most common human hair color, after black. Brown hair is characterized by higher levels of eumelanin and lower levels of pheomelanin. Of the two types of eumelanin (black and brown), brown-haired people have brown eumelanin; they also usually have medium-thick strands of hair. Brown-haired girls or women of European, West Asian or North African descent are often known as brunettes. Chestnut hair is a hair color which is a reddish shade of brown hair. In contrast to auburn hair, the reddish shade of chestnut is darker. Chestnut hair is common among the native peoples of Northern, Central, Western, and Eastern Europe and is also found in Asia Minor, West Asia and North Africa.
Auburn hair ranges along a spectrum of light to dark red-brown shades. The chemicals which cause auburn hair are eumelanin (brown) and pheomelanin (red), with a higher proportion of red-causing pheomelanin than is found in average brown hair. It is most commonly found in individuals of Northern and Western European descent, but is extant in West and Central Asia and North Africa also. It can also be the result of a mutation in the melanocortin 1 receptor gene. [5]
Red hair ranges from light strawberry blond shades to titian, copper, and completely red. Red hair has the highest amounts of pheomelanin, around 67%, and usually low levels of eumelanin. At 1–2% of the west Eurasian population, it is the least common hair color in the world. It is most prominently found in the British Isles and in Udmurtia. Scotland has the highest proportion of redheads; 13 percent of the population has red hair and approximately 40 percent carry the recessive redhead gene. Red hair can also occur in Southern Europe, West Asia, North Africa and Central Asia. [14] [15] [16]
Blond (sometimes blonde for women) hair ranges from pale white (platinum blond) to dark gold blond. Strawberry blond, a mixture of blond and red hair, is a much rarer type containing the most pheomelanin.[ citation needed ] Blond hair can have almost any proportion of pheomelanin and eumelanin, but has only small amounts of both. More pheomelanin creates a more golden or strawberry blond color, and more eumelanin creates an ash or sandy blond color. Blond hair is most commonly found in Northern and Northeastern Europeans and their descendants but can be found spread around most of Europe and also among West Asians and North Africans at lower frequencies. Studies in 2012 showed that naturally blond hair of Melanesians is caused by a recessive mutation in tyrosinase-related protein 1 (TYRP1). In the Solomon Islands, 26% of the population carry the gene; however, it is absent outside of Oceania. [17]
Gray or white hair is not caused by a true gray or white pigment, but is due to a lack of pigmentation and melanin. The clear hairs appear as gray or white because of the way light is reflected from the hairs. Gray hair color typically occurs naturally as people age (see aging or achromotrichia below).
Marie Antoinette syndrome is a proposed phenomenon in which sudden whitening is caused by stress. It has been found that some hairs can become colored again when stress is reduced. [18] [19]
Children born with some hair colors may find it gradually darkens as they grow. Many blond, light brown, or red haired infants experience this. This is caused by genes being turned on and off during early childhood and puberty. [20]
Changes in hair color typically occur naturally as people age, eventually turning the hair gray and then white. This is called achromotrichia. Achromotrichia normally begins in the early to mid-twenties in men and late twenties in women. More than 60 percent of Americans have some gray hair by age 40. The age at which graying begins seems almost entirely due to genetics. Sometimes people are born with gray hair because they inherit the trait. [21]
The order in which graying happens is usually: nose hair, hair on the head, beard, body hair, eyebrows. [22]
Hair color can be changed by a chemical process. Hair coloring is classed as "permanent" or "semi-permanent".
Permanent hair color means that the hair's structure has been chemically altered until it is eventually cut away. This does not mean that the synthetic color will remain permanently. During the process, the natural color is removed, one or more shades, and synthetic color has been put in its place. All pigments wash out of the cuticle. Natural color stays in much longer and artificial will fade the fastest (depending on the color molecules and the form of the dye pigments).
Permanent hair color gives the most flexibility because it can make hair lighter or darker as well as changing tone and color, but there are negatives. Constant (monthly or six-weekly) maintenance is essential to match new hair growing in to the rest of the hair, and to remedy fading. A one-color permanent dye creates a flat, uniform color across the whole head, which can look unnatural and harsh, especially in a fair shade. To combat this, the modern trend is to use multiple colors—usually one color as a base with added highlights or lowlights in other shades.
Semi-permanent color washes out over a period of time—typically four to six weeks, so root regrowth is less noticeable. The final color of each strand is affected by its original color and porosity, so there will be subtle variations in color across the head—more natural and less harsh than a permanent dye. However, this means that gray and white hair will not dye to the same color as the rest of the head (in fact, some white hair will not absorb the color at all). A few gray and white hairs will blend in visually, but semi-permanent dye alone will not usually give the desired result where there is a lot of gray or white hair present. Sometimes a mixture of dyes is used while hair is greying: semi-permanent as a base color, with permanent highlights.
Semi-permanent hair color cannot lighten hair. [23] Hair can only be lightened using chemical lighteners, such as bleach. Bleaching is always permanent because it removes the natural pigment.
"Rinses" are a form of temporary hair color, usually applied to hair during a shampoo and washed out again the next time the hair is washed.
Human skin color ranges from the darkest brown to the lightest hues. Differences in skin color among individuals is caused by variation in pigmentation, which is the result of genetics, exposure to the sun, disorders, or some combination thereof. Differences across populations evolved through natural selection or sexual selection, because of social norms and differences in environment, as well as regulations of the biochemical effects of ultraviolet radiation penetrating the skin.
Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing and painting, brown is usually made by combining the colors orange and black.
Melanin is a family of biomolecules organized as oligomers or polymers, which among other functions provide the pigments of many organisms. Melanin pigments are produced in a specialized group of cells known as melanocytes.
Melanocytes are melanin-producing neural crest-derived cells located in the bottom layer of the skin's epidermis, the middle layer of the eye, the inner ear, vaginal epithelium, meninges, bones, and heart found in many mammals and birds. Melanin is a dark pigment primarily responsible for skin color. Once synthesized, melanin is contained in special organelles called melanosomes which can be transported to nearby keratinocytes to induce pigmentation. Thus darker skin tones have more melanosomes present than lighter skin tones. Functionally, melanin serves as protection against UV radiation. Melanocytes also have a role in the immune system.
Brown hair, also referred to as brunette or brunet, is the second-most common human hair color, after black hair. It varies from light brown to dark hair. It is characterized by higher levels of the dark pigment eumelanin and lower levels of the pale pigment pheomelanin.
Cat coat genetics determine the coloration, pattern, length, and texture of feline fur. The variations among cat coats are physical properties and should not be confused with cat breeds. A cat may display the coat of a certain breed without actually being that breed. For example, a Neva Masquerade could wear point coloration, the stereotypical coat of a Siamese.
Eye color is a polygenic phenotypic trait determined by two factors: the pigmentation of the eye's iris and the frequency-dependence of the scattering of light by the turbid medium in the stroma of the iris.
Bay is a hair coat color of horses, characterized by a reddish-brown or brown body color with a black point coloration on the mane, tail, ear edges, and lower legs. Bay is one of the most common coat colors in many horse breeds.
The silver or silver dapple (Z) gene is a dilution gene that affects the black base coat color and is associated with Multiple Congenital Ocular Abnormalities. It will typically dilute a black mane and tail to a silvery gray or flaxen color, and a black body to a chocolaty brown, sometimes with dapples. It is responsible for a group of coat colors in horses called "silver dapple" in the west, or "taffy" in Australia. The most common colors in this category are black silver and bay silver, referring to the respective underlying coat color.
Equine coat color genetics determine a horse's coat color. Many colors are possible, but all variations are produced by changes in only a few genes. Bay is the most common color of horse, followed by black and chestnut. A change at the agouti locus is capable of turning bay to black, while a mutation at the extension locus can turn bay or black to chestnut.
Black hair is the darkest and most common of all human hair colors globally, due to large populations with this trait. This hair type contains a much more dense quantity of eumelanin pigmentation in comparison to other hair colors, such as brown, blonde and red. In English, various types of black hair are sometimes described as soft-black, raven black, or jet-black. The range of skin colors associated with black hair is vast, ranging from the palest of light skin tones to dark skin. Black-haired humans can have dark or light eyes, but more commonly dark.
Greying of hair, also known as greying, canities, or achromotrichia, is the progressive loss of pigmentation in the hair, eventually turning the hair grey or white which typically occurs naturally as people age.
The genetic basis of coat colour in the Labrador Retriever has been found to depend on several distinct genes. The interplay among these genes is used as an example of epistasis.
Light skin is a human skin color that has a low level of eumelanin pigmentation as an adaptation to environments of low UV radiation. Due to migrations of people in recent centuries, light-skinned populations today are found all over the world. Light skin is most commonly found amongst the native populations of Europe, East Asia, West Asia, Central Asia, Siberia, and North Africa as measured through skin reflectance. People with light skin pigmentation are often referred to as "white" although these usages can be ambiguous in some countries where they are used to refer specifically to certain ethnic groups or populations.
Amelanism is a pigmentation abnormality characterized by the lack of pigments called melanins, commonly associated with a genetic loss of tyrosinase function. Amelanism can affect fish, amphibians, reptiles, birds, and mammals including humans. The appearance of an amelanistic animal depends on the remaining non-melanin pigments. The opposite of amelanism is melanism, a higher percentage of melanin.
A melanistic mask is a dog coat pattern that gives the appearance of a mask on the dog's face. The hairs on the muzzle, and sometimes entire face or ears, are colored by eumelanin instead of pheomelanin pigment. Eumelanin is typically black, but may instead be brown, dark gray, or light gray-brown. Pheomelanin ranges in color from pale cream to mahogany. The trait is caused by M264V (EM), a completely dominant allele (form) of the melanocortin 1 receptor gene.
Dark skin is a type of human skin color that is rich in melanin pigments. People with dark skin are often referred to as black people, although this usage can be ambiguous in some countries where it is also used to specifically refer to different ethnic groups or populations.
Albinism is the congenital absence of melanin in an animal or plant resulting in white hair, feathers, scales and skin and reddish pink or blue eyes. Individuals with the condition are referred to as albinos.
Dogs have a wide range of coat colors, patterns, textures and lengths. Dog coat color is governed by how genes are passed from dogs to their puppies and how those genes are expressed in each dog. Dogs have about 19,000 genes in their genome but only a handful affect the physical variations in their coats. Most genes come in pairs, one being from the dog's mother and one being from its father. Genes of interest have more than one expression of an allele. Usually only one, or a small number of alleles exist for each gene. In any one gene locus a dog will either be homozygous where the gene is made of two identical alleles or heterozygous where the gene is made of two different alleles.
The agouti gene, the Agouti-signaling protein (ASIP) is responsible for variations in color in many species. Agouti works with extension to regulate the color of melanin which is produced in hairs. The agouti protein causes red to yellow pheomelanin to be produced, while the competing molecule α-MSH signals production of brown to black eumelanin. In wildtype mice, alternating cycles of agouti and α-MSH production cause agouti coloration. Each hair has bands of yellow which grew during agouti production, and black which grew during α-MSH production. Wildtype mice also have light-colored bellies. The hairs there are a creamy color the whole length because the agouti protein was produced the whole time the hairs were growing.