Kokomopteroidea

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Kokomopteroidea
Temporal range: Early Silurian - Late Devonian, 443.7–376.1  Ma
The Eurypterida of New York plate 54.jpg
Reconstruction of Kokomopterus , a kokomopterid.
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Subphylum: Chelicerata
Order: Eurypterida
Suborder: Stylonurina
Superfamily: Kokomopteroidea
Kjellesvig-Waering, 1966
Families

Kokomopteroidea is an extinct superfamily of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". It is one of four superfamilies classified as part of the suborder Stylonurina. Kokomopteroids have been recovered from deposits of Early Silurian to Late Devonian age in the United States and the United Kingdom. [1]

Contents

Description

Kokomopteroids are stylonurines with a spiniferous appendage V and a posterior notch on the metastoma. The superfamily forms two distinct clades; the Kokomopteridae (including the genera Kokomopterus and Lamontopterus ) and the Hardieopteridae (including the genera Hardieopterus , Tarsopterella and Hallipterus ). [2]

The Kokomopteroidea retains primitive Hughmilleria -type prosomal appendages for unsuitable raking through the bottom sediments of marine environments. As such, the members of the superfamily were likely scavengers. The Hardieopteridae display some sweep-feeding characteristics similarly to members of the superfamilies Hibbertopteroidea and Stylonuroidea, including a broadening of the metastoma and the spines of the anterior prosomal appendages. Hardieopterids may even have been benthic bottomdwellers partially living buried in the substrate. [2]

Systematics and genera

The Kokomopteroids are eurypterids classified as part of the suborder Stylonurina. They are a sister group to the Hibbertopteroidea, united by a median ridge on the carapace between the lateral eyes and a distal thickening to the podomeres of the prosomal appendages. The primitive hibbertopteroid Drepanopterus is a sister taxon to all other hibbertopteroids and shares certain characteristics with the Kokomopteroidea (such as a clavate telson). [2]

Kokomopteroidea contains two families, the Kokomopteridae and Hardieopteridae. The Kokomopteridae is defined as kokomopteroids with undifferentiated opisthosoma with a marginal rim and the appendages II-V being spiniferous as in Lamontopterus , with appendage VI not being spiniferous, as in Kokomopterus . The Hardieopteridae is defined as kokomopteroids with lateral pleurae on the metasoma and pretelson, large lunate scales on the posterior carapace margin and a clavate telson. The opisthosoma shows both second and fourth order differentiation into a mesosoma and metasoma in the Hardieopteridae, with appendage V being spiniferous and VI being non-spiniferous as in Hardieopterus. The Kokomopteridae is only known from the Silurian, the temporal range of the Hardieopteridae extends further to the Late Devonian. [2]

Superfamily KokomopteroideaKjellesvig-Waering, 1966

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Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus Eurypterus accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Triassic extinction event 251.9 million years ago.

<span class="mw-page-title-main">Stylonurina</span> Extinct suborder of arthropods

Stylonurina is one of two suborders of eurypterids, a group of extinct arthropods commonly known as "sea scorpions". Members of the suborder are collectively and informally known as "stylonurine eurypterids" or "stylonurines". They are known from deposits primarily in Europe and North America, but also in Siberia.

<i>Slimonia</i> Extinct genus of arthropods

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<i>Hughmilleria</i> Genus of extinct arthropods

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<i>Drepanopterus</i> Extinct genus of sea scorpions

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<i>Nanahughmilleria</i> Extinct genus of arthropods

Nanahughmilleria is a genus of eurypterid, an extinct group of aquatic arthropods. Fossils of Nanahughmilleria have been discovered in deposits of Devonian and Silurian age in the United States, Norway, Russia, England and Scotland, and have been referred to several different species.

<i>Onychopterella</i> Extinct genus of arthropods

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<i>Erettopterus</i> Extinct genus of arthropods

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<span class="mw-page-title-main">Pterygotidae</span> Extinct family of eurypterids

Pterygotidae is a family of eurypterids, an extinct group of aquatic arthropods. They were members of the superfamily Pterygotioidea. Pterygotids were the largest known arthropods to have ever lived with some members of the family, such as Jaekelopterus and Acutiramus, exceeding 2 metres (6.6 ft) in length. Their fossilized remains have been recovered in deposits ranging in age from 428 to 372 million years old.

<span class="mw-page-title-main">Stylonuridae</span> Extinct family of arthropods

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<span class="mw-page-title-main">Stylonuroidea</span> Extinct superfamily of arthropods

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<span class="mw-page-title-main">Parastylonuridae</span> Extinct family of arthropods

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<span class="mw-page-title-main">Kokomopteridae</span> Extinct family of arthropods

The Kokomopteridae are a family of eurypterids, an extinct group of chelicerate arthropods commonly known as "sea scorpions". The family is one of two families contained in the superfamily Kokomopteroidea, which in turn is one of four superfamilies classified as part of the suborder Stylonurina.

<span class="mw-page-title-main">Hardieopteridae</span> Extinct family of arthropods

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<span class="mw-page-title-main">Adelophthalmidae</span> Family of eurypterids, an extinct group of aquatic arthropods

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<span class="mw-page-title-main">Hughmilleriidae</span> Extinct family of eurypterids

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References

  1. "Fossilworks: Kokomopteroidea". fossilworks.org. Retrieved 17 December 2021.
  2. 1 2 3 4 James C. Lamsdell, Simon J. Braddy & O. Erik Tetlie (2010). "The systematics and phylogeny of the Stylonurina (Arthropoda: Chelicerata: Eurypterida)". Journal of Systematic Palaeontology . 8 (1): 49–61. Bibcode:2010JSPal...8...49L. doi: 10.1080/14772011003603564 .