Manipulator Temporal range: Kimmeridgian to Cenomanian, | |
---|---|
Fossil specimen | |
Reconstruction | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Blattodea |
Superfamily: | Corydioidea |
Family: | † Manipulatoridae Vršanský & Bechly 2015 [1] |
Genus: | † Manipulator Vršanský & Bechly 2015 [1] |
Type species | |
Manipulator modificaputis Vršanský & Bechly 2015 | |
Other species | |
Manipulator olimVršanský, 2024 |
Manipulator is an extinct cockroach genus including species which lived from the Late Jurassic to the Late Cretaceous. The type species is Manipulator modificaputis, described on the basis of the holotype specimen which is fossilized in a 100-million-year-old piece of Burmese amber, found in a quarry of volcanoclastic mudstone (a sedimentary rock) at Noije Bum in the Hukawng Valley in Myanmar. M. modificaputis was described by Peter Vršanský, of the Geological Institute SAS of Bratislava, and by Günter Bechly, of the Staatliches Museum für Naturkunde in Stuttgart. [1]
The cockroach was found to have an elongated neck, a freely rotating head and unexpectedly long legs, which are indicative of a predatory lifestyle. [2] However, as of 2022, it is thought that M. modificaputis tended to live around and feed on flowers. [3] In addition, this insect has some characteristics superficially similar to mantises, however differ from other many characters. [3] The insect body is 9.3 millimetres (0.37 in) long with 14.7 millimetres (0.58 in) long forewings for male and 10.9 millimetres (0.43 in) body length, 13.4 millimetres (0.53 in) forewing length for female. [3]
The authors erected a new family, Manipulatoridae, after examining the specimen on the basis of "the unique habitus with numerous autapomorphies along with several plesiomorphies." [4] Other specimens including that of a juvenile were discovered from the Myanmar amber mines [4] and described in 2022. [3]
This species was found along with dozens of other extinct species of insects fully preserved in amber, making Noije Bum in the Hukawng Valley region one of the most important regions for amber fossils containing fully preserved insects. [5]
This species belonged to the invertebrate fauna of the ancient amber forest of the Myanmar region. [1]
Vršanský (2024) described another species belonging to the genus Manipulator, Manipulator olim from the Upper Jurassic (Kimmeridgian) Karabastau Formation (Kazakhstan). [6]
Manipulator was the only member of the family Manipulatoridae until 2022, when the related genus Manipulatoides was described from the same deposit as the type species. [3]
Corydiidae, previously known as Polyphagidae, is a family of the order Blattodea (cockroaches). Many are known as sand cockroaches. The family is divided into five subfamilies, comprising some 40 genera. One prominent species is the desert cockroach, Arenivaga investigata.
Proraphidia is a genus of snakefly in the extinct family Mesoraphidiidae. The genus currently contains three species; Proraphidia gomezi from the La Pedrera de Rúbies Formation in Spain, Proraphidia hopkinsi from the Weald Clay in England, and the type species Proraphidia turkestanica from Kazakhstan. The genus was first described by O. M. Martynova in 1941 with the publication of P. turkestanica from Jurassic deposits of the Karabastau Formation in Karatau, Kazakhstan.
Myanmymar is an extinct genus of fairyfly preserved in Burmese amber from Myanmar. It has only one species, Myanmymar aresconoides. It is dated to the earliest part of the Cenomanian stage of the Late Cretaceous, around 99 million years old. As of 2011, it is the oldest known fossil mymarid.
Mesoraphidiidae is an extinct family of snakeflies in the suborder Raphidiomorpha. The family lived from the Late Jurassic through the Late Cretaceous and is known from twenty-five genera. Mesoraphidiids have been found as both compression fossils and as inclusions in amber. The family was first proposed in 1925 by the Russian paleoentomologist Andrey Vasilyevich Martynov based on Upper Jurassic fossils recovered in Kazakhstan. The family was expanded in 2002 by the synonymizing of several other proposed snakefly families. The family was divided into three subfamilies and one tribe in a 2011 paper, further clarifying the relationships of the included genera.
The Stenophlebiidae is an extinct family of medium-sized to large fossil odonates from the Upper Jurassic and Cretaceous period that belongs to the damsel-dragonfly grade ("anisozygopteres") within the stem group of Anisoptera. They are characterized by their long and slender wings, and the transverse shape of the discoidal triangles in their wing venation.
Lebanoraphidia is an extinct genus of snakefly in the family Mesoraphidiidae. The genus is solely known from Cretaceous, Upper Neocomian, fossil amber found in Lebanon. Currently the genus is composed of a single species Lebanoraphidia nana.
Chresmoda is an extinct genus of insects within the family Chresmodidae.
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly arthropods including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to the potential role of the amber trade in funding internal conflict in Myanmar and hazardous working conditions in the mines where it is collected.
Rafaelnymphes is an extinct genus of lacewing in the family Nymphidae known from a fossil found in South America. The genus contains a single species, Rafaelnymphes cratoensis.
Alienopterus brachyelytrus is an extinct insect described from a 99 million year old fossil found in Burmese amber from the Hukawng Valley of Myanmar. It was the first known member of the order Alienoptera until 2018, when the second and third members of the order, Caputoraptor elegans, and Alienopterella stigmatica were described. A. brachyelytrus has characters that are shared with cockroaches and mantids and is thought to represent either the sister taxon, or an ancestor to mantids.
Alienopteridae is an extinct family of dictyopterans, known from the Early Cretaceous to the Eocene epoch. They are noted for their unusual combination of features not found in other dictyopterans.
Aethiocarenus is an extinct genus of insects which has a single species Aethiocarenus burmanicus described from a 98.79 ±0.62 million year old fossil found in Burmese amber from the Hukawng Valley of Myanmar. The insect is unusual due to the vertex of the triangular head being attached to the pronotum as opposed to the hypotenuse. When first described Aethiocarenus was placed as the sole member of the family Aethiocarenidae and order Aethiocarenodea. However, Aethiocarenus was later considered to be a nymph of Alienopterus. Vršanský et al. (2018) considered Aethiocarenus to be an alienopterid nymph, but considered it distinct from other members of this group and deserving a separate genus rank.
Caputoraptor is an extinct genus of insect from the Cenomanian aged Burmese amber containing two species: the type species Caputoraptor elegans, as well as Caputoraptor vidit. It is part of the extinct order Alienoptera. C. elegans is notable for the presence of a scissor like mechanism consisting of a straight edge on the back of the head and corresponding serrated edges on the first thoracic segment, these were initially suggested to be used by the female to grasp the male during mating, but the structure is not sexually dimorphic, so a use to hold prey was subsequently suggested. Its morphology suggests a predatory habit inhabiting shrubs and trees. In 2020 a Caputoraptor elegans nymph was described that was in the process of being predated upon by a Ceratomyrmex hell ant.
2019 in paleoentomology is a list of new fossil insect taxa that were described during the year 2019, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
2017 in paleoentomology is a list of new fossil insect taxa that were described during the year 2017, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
Permopsocida is an extinct order of insects known from the Carboniferous (Moscovian) to the Mid-Cretaceous. It is part of Paraneoptera, alongside bark lice, bugs and thrips. Within Paraneoptera it is considered to be closer to the clade containing bugs and thrips rather than bark lice, with an estimated divergence during the Late Carboniferous. The group was first named as a suborder by Robert John Tillyard in 1926, and was raised to a full order by Huang et al. in 2016. It is currently divided up into three families, Psocidiidae which is known from the Late Carboniferous to Liassic. Permopsocidae which is only known from the Permian, and Archipsyllidae, which is known from the Late Triassic to mid-Cretaceous (Cenomanian). While most members of the group are known from compression fossils, several members of Archipsyllidae are 3 dimensionally preserved in Burmese amber, which has helped clarify the morphology and phylogenetic position of the group. The morphology of the mouthparts suggests that they were capable of suction feeding and chewing, with preserved angiosperm pollen grains in the gut of Psocorrhyncha suggesting that at least some members of the group were pollenivorous.
Blattulidae is an extinct family of cockroaches known from the Triassic to the Late Cretaceous. Their distinguishing characteristics include "forewing has long Sc, regular venation with distinct intercalaries and hindwing has simple CuP, branched A1." Due to the poor ability of forewing venation to correctly classify modern cockroaches to extensive homoplasy, the value of this family as a taxonomic unit has been strongly questioned, with some authors considering the family a nomen dubium.
Mesoblattinidae is an extinct, problematic family of cockroaches known from the Mesozoic. It was formerly considered a wastebasket taxon for Mesozoic cockroaches, but the family has subsequently been better defined, with many taxa transferred to Caloblattinidae. It is considered to have close affinities with Blattidae and Ectobiidae, as well as possibly Blaberidae. The family first appeared by the Early Jurassic. They are considered to represent amongst the oldest groups of modern cockroaches, and like them are thought to have laid oothecae. Due to the poor ability of forewing venation to correctly classify cockroaches to extensive homoplasy, the value of this family as a taxonomic unit has been strongly questioned, with some authors considering the family a nomen dubium.
Liberiblattinidae is an extinct family of cockroaches known from the Jurassic to Cretaceous. Some taxa, like Cryptoblatta and Hydrokhoohydra, are suggested to be semiaquatic. Spongistoma is suggested to be a nectarivore due to its unique sucking/sponging "proboscis" mouthparts. Some authors have suggested that the family is ancestral to Mantodea.
Paleotriatoma metaxytaxa is a species of fossil insect belonging to the subfamily Triatominae of the family Reduviidae. Living kissing bugs are blood-sucking insects responsible for the transmission of Chagas disease. Chagas is a parasitic disease affecting millions of people mainly in South America, Central America and Mexico.