Masillaraptor

Last updated

Masillaraptor
Temporal range: Middle Eocene
Masillaraptor restoration 2.jpeg
Life restoration based on modern falconiformes
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Falconiformes
Family: Masillaraptoridae
Genus: Masillaraptor
Mayr, 2006
Species:
M. parvunguis
Binomial name
Masillaraptor parvunguis
Mayr, 2006

Masillaraptor is an extinct genus of masillaraptorid, a group of primitive falconiforms, from the Middle Eocene Messel Pit, Germany. It is a long-legged relative of the living falcons. [1] [2]

Contents

Known species

Only one species of Masillaraptor is known: M. parvunguis

Etymology

Masillaraptor comes from the Latin word Masilla, which is the old name for the town of Messel, and raptor is a New Latin suffix used to indicate a predator (from rapere, to catch) and in English it means bird of prey.

Specific epithet parvunguis is also Latin, coming from the word parvus which means small and feeble, while unguis means claw.

The name refers to the fact that the specimen's claws are small in comparison to those of other raptors.

Characteristics

The genus Masillaraptor is different from all other known avian taxa. It possesses a combination of characters that distinguishes it from all others.

1. The beak is almost as long as the cranium itself, with equal height over much of its length and a straight dorsal ridge. The beak curves just before its tip, restricting the nasal openings to the rear half of the beak.

2. The tibiotarsus is the longest bone in the leg.

3. On the second toe the first phalanx is shortened, whereas on the fourth toe the second and third phalanges are shortened.

4. The claws of Masillaraptor are small and weak compared to other falconiform birds with abbreviated pedal phalanges.

Characters (1) and (3) are derived within neornithine birds and also found in modern Accipitres, from which Masillaraptor is, however, distinguished in character (4).(Mayr, 2006.)

Fossil specimens

There are two specimens of Masillaraptor. Only one specimen was referenced for classification because the other is housed in an unknown private collection. Both specimens are a slab of rock containing a nearly complete, articulate but poorly preserved skeleton. The specimens are both believed to be adult members of the species. Both specimens were discovered in the Messel pit, an old shale mine known for the extremely well preserved fossils that have been discovered there.

Classification

Masillaraptor represents one of two members of Masillaraptoridae within the Falconiformes. The cladogram below displays the results of the phylogenetic analysis by Mayr & Kitchener (2022): [3]

Chauna torquata (southern screamer)
Chauna torquata BYN.jpg
Australaves
Falconiformes

Caracara plancus (crested caracara) Caracara1.png

Masillaraptoridae

Masillaraptor parvunguis Masillaraptor restoration.jpeg

Danielsraptor phorusrhacoides

Cariamiformes
Bathornis spp.
Bathornis grallator restoration.jpg

Dynamopterus spp.

Cariama cristata (red-legged seriema) Cariama cristata 1838 white background.jpg

Phorusrhacidae  (terror birds)

Llallawavis scagliai Llallawavis scagliai.jpg

Psilopterus lemoinei
Psilopterus.jpg

Patagornis marshi Patagornis marshi.png

Related Research Articles

<span class="mw-page-title-main">Falcon</span> Birds of prey in the genus Falco

Falcons are birds of prey in the genus Falco, which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene.

<span class="mw-page-title-main">Seriema</span> Family of birds

The seriemas are the sole living members of the small bird family Cariamidae, which is also the only surviving lineage of the order Cariamiformes. Once believed to be related to cranes, they have been placed near the falcons, parrots and passerines, as well as the extinct Phorusrhacidae. The seriemas are large, long-legged territorial birds that range from 70–90 cm (28–35 in) in length. They live in grasslands, savanna, dry woodland and open forests of Brazil, Bolivia, Argentina, Paraguay and Uruguay. There are two species of seriemas, the red-legged seriema and the black-legged seriema. Names for these birds in the Tupian languages are variously spelled as siriema, sariama, and çariama, and mean "crested".

<span class="mw-page-title-main">Coraciidae</span> Family of birds

Coraciidae is a family of Old World birds, which are known as rollers because of the aerial acrobatics some of these birds perform during courtship or territorial flights. Rollers resemble crows in size and build, and share the colourful appearance of kingfishers and bee-eaters, blues and pinkish or cinnamon browns predominating. The two inner front toes are connected, but not the outer one.

<span class="mw-page-title-main">Phorusrhacidae</span> Extinct family of flightless birds

Phorusrhacids, colloquially known as terror birds, are an extinct family of large carnivorous flightless birds that were among the largest apex predators in South America during the Cenozoic era; their conventionally accepted temporal range covers from 53 to 0.1 million years (Ma) ago.

<span class="mw-page-title-main">Plotopteridae</span> Family of sea birds

Plotopteridae is an extinct family of flightless seabirds with uncertain placement, generally considered as member of order Suliformes. They exhibited remarkable convergent evolution with the penguins, particularly with the now extinct giant penguins. That they lived in the North Pacific, the other side of the world from the penguins, has led to them being described at times as the Northern Hemisphere's penguins, though they were not closely related. More recent studies have shown, however, that the shoulder-girdle, forelimb and sternum of plotopterids differ significantly from those of penguins, so comparisons in terms of function may not be entirely accurate. Plotopterids are regarded as closely related to Anhingidae (darters) and Phalacrocoracidae (cormorants). On the other hand, there is a theory that this group may have a common ancestor with penguins due to the similarity of forelimb and brain morphology. However, the endocast morphology of stem group Sphenisciformes differs from both Plotopteridae and modern penguins.

<i>Phorusrhacos</i> Extinct genus of birds

Phorusrhacos is an extinct genus of giant flightless terror birds that inhabited South America during the Miocene epoch. Phorusrhacos was one of the dominant land predators in South America at the time it existed. It is thought to have lived in woodlands and grasslands.

Eurofluvioviridavis is a genus of extinct primitive birds from the Middle Eocene Messel Pit, Germany. It contains a single species, Eurofluvioviridavis robustipes. It is related to Avolatavis and Vastanavis, other members of the family Vastanavidae.

<span class="mw-page-title-main">Falconiformes</span> Order of birds

The order Falconiformes is represented by the extant family Falconidae and a handful of enigmatic Paleogene species. Traditionally, the other bird of prey families Cathartidae, Sagittariidae (secretarybird), Pandionidae (ospreys), Accipitridae (hawks) were classified in Falconiformes. A variety of comparative genome analysis published since 2008, however, found that falcons are part of a clade of birds called Australaves, which also includes seriemas, parrots and passerines. Within Australaves falcons are more closely related to the parrot-passerine clade (Psittacopasserae), which together they form the clade Eufalconimorphae. The hawks and vultures occupy a basal branch in the clade Afroaves in their own clade Accipitrimorphae, closer to owls and woodpeckers.

<span class="mw-page-title-main">Bohaiornithidae</span> Extinct family of dinosaurs

Bohaiornithidae is a group of early predatory enantiornitheans from the early Cretaceous Period of China. All known specimens come from the Jiufotang Formation and Yixian Formation, dating to the early Aptian age, 125–120 million years ago. Bohaiornithidae was first coined as a family of enantiornithean birds by Wang and colleagues in 2014. They defined it as the natural group formed by all descendants of the common ancestor of the type species, Bohaiornis guoi, and Shenqiornis mengi.

<span class="mw-page-title-main">Messelasturidae</span> Extinct family of birds

Messelasturidae is an extinct family of birds known from the Eocene of North America and Europe. Their morphology is a mosaic that in some aspects are very similar to modern hawks and falcons, but in others are more similar to parrots. Initially interpreted as stem-owls, more recent studies have suggested a closer relationship to parrots and passerines. Their ecology is enigmatic.

Vastanavidae is an extinct family of birds related to parrots and passerine birds. They are known from fossils from Eocene sites in India, Europe, and North America. The vastanavids resemble parrots and the extinct parrot relative Quercypsitta in their morphology, including the partially zygodactyl foot, in which two toes could face opposite the other two.

<span class="mw-page-title-main">Halcyornithidae</span> Extinct family of birds

Halcyornithidae is an extinct family of telluravian birds thought to be related to the Psittaciformes (parrots), Passeriformes (songbirds), and to the extinct Messelasturidae. Halcyornithids have been found in various Eocene formations in Europe and North America. Widespread and diverse in the Early Eocene of North America and Europe, halcyornithids are not found in locales later than the Middle Eocene. Halcyornithids were small, arboreal birds with zygodactyl feet, with two toes facing forwards and two facing back, a trait shared with other tree-dwelling families of Eocene birds like the Zygodactylidae and the messelasturids. The skull of halcyornithids features a ridge of bone above the eye called the supraorbital process, similar to birds of prey. The relationships of the halcyornithids to other birds remain uncertain. Halcyornithids have been proposed as relatives to owls and as a lineage closer to parrots than to songbirds. Most recently, halcyornithids have been identified as the sister group of the clade including parrots and songbirds. It is also possible that Halcyornithidae is paraphyletic with respect to the Messelasturidae.

Oligocolius is an unusual genus of extinct mousebird from the early to late Oligocene epoch of Germany. Oligocolius is known from two species, the type species O. brevitarsus and O. psittacocephalon. Oligocolius is a member of the family of modern mousebirds (Coliidae) and is broadly similar to them in shape. However, unlike modern mouse birds the skull and beak of Oligocolius closely resembles those of parrots, and appears to be specialised for a distinct lifestyle not found in living mousebirds.

<i>Eocoracias</i> Extinct genus of birds

Eocoracias is an extinct genus of bird related to modern rollers and other Coraciiformes such as kingfishers, bee-eaters, motmots, and todies. It contains one species, Eocoracias brachyptera, and it lived approximately 47 million years ago based on dating of the fossil site. It is known for a specimen having preserved non-iridescent structural coloration on its feathers, previously unknown in fossil birds. Fossils have been found at the Messel Pit in Germany.

This is an overview of the paleofauna of the Eocene Messel Formation as explored by the Messel Pit excavations in Germany. A former quarry and now UNESCO World Heritage Site, the Messel Formation preserves what once were a series of anoxic lakes surrounded by a sub-tropical rainforest during the Middle Eocene, approximately 47 Ma.

<i>Scaniacypselus</i> Extinct genus of bird

Scaniacypselus is an extinct genus of basal swifts from the Eocene of Denmark, Germany and France. Many well preserved specimens still showing plumage were discovered in the Messel Pit near Darmstadt, Germany. Species of Scaniacypselus had relatively broader wings than modern swifts and hindlimbs better adapted to perching on tree-branches, indicating that the bird was not as aerial as its extant relatives and likely nested in trees like hummingbirds and treeswifts. Two species are recognized, S. wardi and S. szarskii.

<span class="mw-page-title-main">Masillaraptoridae</span> Extinct family of birds

Masillaraptoridae is an extinct family of stem-group falconiform birds from the Eocene of Europe. They are noted for their relatively long legs. Two genera have been named: Danielsraptor, from the London Clay of England, and Masillaraptor, from Messel Pit in Germany.

Danielsraptor is an extinct genus of masillaraptorid bird from the Early Eocene (Ypresian) Walton Member of the London Clay Formation in Essex, United Kingdom. The genus contains a single species, D. phorusrhacoides, known from a partial skeleton.

<i>Waltonavis</i> Extinct genus of birds

Waltonavis is an extinct genus of potentially leptosomiform bird from the Early Eocene London Clay Formation of Essex, United Kingdom. The genus contains two species: W. paraleptosomus and W. danielsi, both known from partial skeletons.

<i>Serudaptus</i> Extinct genus of birds

Serudaptus is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany. The genus contains one species, Serudaptus pohli, and is known for long, raptorial claws on its zygodactyl feet.

References

  1. Mayr, G. (2006) A new raptorial bird from the Middle Eocene of Messel, Germany. Historical Biology, 18(2): 95–102
  2. Mayr G, Kitchener AC (2022). "New fossils from the London Clay show that the Eocene Masillaraptoridae are stem group representatives of falcons (Aves, Falconiformes)". Journal of Vertebrate Paleontology. 41 (6): e2083515. doi: 10.1080/02724634.2021.2083515 . S2CID   250402777.
  3. Mayr, Gerald; Kitchener, Andrew C. (2022-07-08). "New fossils from the London Clay show that the Eocene Masillaraptoridae are stem group representatives of falcons (Aves, Falconiformes)". Journal of Vertebrate Paleontology. 41 (6): e2083515. doi: 10.1080/02724634.2021.2083515 . ISSN   0272-4634. S2CID   250402777.