Oryctorhynchus Temporal range: Late Triassic, ~ | |
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Skeletal reconstruction of O. bairdi | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Clade: | Archosauromorpha |
Order: | † Rhynchosauria |
Family: | † Rhynchosauridae |
Subfamily: | † Hyperodapedontinae |
Genus: | † Oryctorhynchus Sues, Fitch & Whatley, 2020 |
Type species | |
†Oryctorhynchus bairdi Sues, Fitch & Whatley, 2020 |
Oryctorhynchus is an extinct genus of rhynchosaur from the Late Triassic (Carnian-Norian)-aged Wolfville Formation of Nova Scotia, Canada that may have been the same animal as Beesiiwo . [1] The type species, O. bairdi, was named and described in 2020. [2] It was originally seen as a species of Hyperodapedon until 2020.
The holotype was discovered in the Wolfville Formation by Donald Baird; its earliest known mention is by Baird (1963). [3] Shortly after, it was informally named the "Nova Scotia Hyperodapedon " (H. sp.) by Robin Whatley in a 1984 paper published by J. A. Hopson. [4] It was briefly described by Michael Benton (1983) also as a species of Hyperodapedon. [5] It was then assigned to cf. "Hyperodapedon" sanjuanensis by Lucas etal., (2002). [6] The genus was not recognised as a distinct taxon until it was named in 2020. [2]
The holotype, NSM018GF009.012, consists of a partial jaw and several skull fragments including the rostrum and skull roof. [2] [5]
Fitch et al. (2023) state that specimen NSM018GFF009.003 has "No unique support for [being] Oryctorhynchus bairdi, and [they] do not consider it a part of O . bairdi. These attributes better align with those found in Beesiiwo cooowuse ... [they] suggest it is either a close relative of Beesiiwo or a member of such." [1]
The genus name consists of the orycto prefix, which means burrow, and the rhynchus suffix, meaning snout; the full genus name means burrowed snout. The epithet honours David Baird, for his work on Triassic tetrapods from Nova Scotia. [2]
Sues et al. (2020) placed Oryctorhynchus as the sister species to Hyperodapedon and an unnamed hyperodapedontine taxon from Wyoming. [2]
Oryctorhynchus is from the Wolfville Formation (Upper Wolfville Member; Fundy Basin), which probably corresponds to the Popo Agie Formation. The age of the Upper Wolfville Member is unclear; it either dates from the latest Carnian? - earliest Norian? or the late Carnian (~230 Ma). [7]
It would have coexisted with Acadiella , [8] Arctotraversodon , [9] Arctosuchus buceros (?) , [10] Haligonia , [8] Scoloparia [8] and Teraterpeton . [11]
The Carnian is the lowermost stage of the Upper Triassic Series. It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by the Norian. Its boundaries are not characterized by major extinctions or biotic turnovers, but a climatic event occurred during the Carnian and seems to be associated with important extinctions or biotic radiations. Another extinction occurred at the Carnian-Norian boundary, ending the Carnian age.
Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaurs first appeared in the Early Triassic, reaching their broadest abundance and a global distribution during the Carnian stage of the Late Triassic.
Hyperodapedon is an extinct genus of rhynchosaur reptiles which lived during Late Triassic period. Like other rhynchosaurs, it was an heavily built archosauromorph, distantly related to archosaurs such as crocodilians and dinosaurs. Hyperodapedon in particular was part of the subfamily Hyperodapedontinae, a specialized rhynchosaurian subgroup with broad skulls, beaked snouts, and crushing tooth plates on the roof of the mouth.
Paleorhinus is an extinct genus of widespread basal phytosaur known from the Late Triassic. The genus was named in 1904 based on the type species Paleorhinus bransoni, which is known from Wyoming and Texas in the United States. Another valid species, Paleorhinus angustifrons from Bavaria, Germany, is also commonly referred to the genus. Paleorhinus had a length of about 2.5 meters.
The Ischigualasto Formation is a Late Triassic geological formation in the Ischigualasto-Villa Unión Basin of southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation dates to the late Carnian and early Norian stages of the Late Triassic, according to radiometric dating of ash beds.
Jachaleria was a dicynodont herbivore that lived from the Ladinian to Norian stages of the Middle to Late Triassic, from approximately 240 to 220 million years ago. Jachaleria was one of the last representatives of the dicynodonts, occurring in Argentina and Brazil. It lacked teeth, much like Stahleckeria, but was closer in size to Dinodontosaurus.
Angistorhinus is an extinct genus of phytosaur known from the Late Triassic period of Texas and Wyoming, United States. It was first named by Mehl in 1913 and the type species is Angistorhinus grandis. Other species from Texas and Wyoming, A. alticephalus, A. gracilis and A. maximus, are cospecific with the type species. Angistorhinus is known from the holotype UC 631, partial skull and lower jaws recovered from the Popo Agie Formation, Chugwater Group, Wyoming and from the associated paratype UM 531, a partial skull, TMM 31098-1, skull and lower jaws and ROM 7977, partial skull and lower jaws, recovered from the 'Pre-Tecovas Horizon' in the Dockum Group, Texas. A possible second species, A. talainti is known from the Triassic of Morocco. In 1995, Long and Murry created the new combination, Angistorhinus megalodon by synonymy for Brachysuchus. Hungerbühler and Sues (2001) hypothesised that Angistorhinus is a junior synonym of Rutiodon. However, in 2010 Michelle R. Stocker retained the validity of Brachysuchus and of A. grandis.
Heptasuchus is an extinct genus of loricatan pseudosuchian known from the Middle or Late Triassic upper Chugwater Group of Wyoming, United States. It contains a single species, Heptasuchus clarki, the first formally recognized "rauisuchian" or loricatan pseudosuchian from North America.
The Popo Agie Formation is a Triassic geologic formation that crops out in western Wyoming, western Colorado, and Utah. It was deposited during the Late Triassic in fluvial (river) and lacustrine (lake) environments that existed across much of what is now the American southwest. Fragmentary fossils of prehistoric reptiles and amphibians, including pseudosuchian reptiles and temnospondyl amphibians, have been discovered in the Popo Agie Formation. Dinosaur remains are also among the fossils that have been recovered from the formation, although none have yet been referred to a specific genus.
The Tiki Formation is a Late Triassic geologic formation in Madhya Pradesh, northern India. Dinosaur remains are among the fossils that have been recovered from the formation, although none have yet been referred to a specific genus. Phytosaur remains attributable to the genus Volcanosuchus have also been found in the Tiki Formation.
The Wolfville Formation is a Triassic geologic formation of Nova Scotia. The formation is of Carnian to early Norian age. Fossils of small land vertebrates have been found in the formation, including procolophonid and early archosauromorph reptiles and cynodonts. Dinosaur remains are among the fossils that have been recovered from the formation, although none have yet been referred to a specific genus.
Isalorhynchus is an extinct genus of hyperodapedontine rhynchosaur from the late Triassic period of Toliara Province, southwestern Madagascar. It is known from the holotype MDE-R18, a nearly complete maxilla and from other specimens from the same locality, Malio River area. It was found in the Makay Formation of the Morondava Basin. It was first named by Eric Buffetaut in 1983 and the type species is Isalorhynchus genovefae. The majority of Isalorhynchus specimens are isolated jaw bones, but two nearly complete skeletons were found in 1998. Langer et al., 2000 concluded that Isalorhynchus is a synonym of Hyperodapedon and referred it to a new species of Hyperodapedon. Whatley, 2005 retained this genus as valid with a description of new materials in her PhD thesis. Montefeltro et al., 2010 and Langer et al., 2010 accepted Isalorhynchus as valid genus.
Sanjuansaurus is a genus of herrerasaurid dinosaur from the Late Triassic (Carnian) Ischigualasto Formation of the Ischigualasto-Villa Unión Basin in northwestern Argentina.
Hyperodapedontinae is a subfamily of rhynchosaurs within the family Rhynchosauridae. Fossils have been found from Argentina, Brazil, Canada, India, Madagascar, Scotland, Tanzania, United States and Zimbabwe.
Macrocephalosaurus is a genus of rhynchosaurs from the Late Triassic period of southern Brazil. Although usually synonymized with Hyperodapedon, recent cladistic analysis has called this synonymy into question.
Teraterpeton is an extinct genus of trilophosaurid archosauromorphs. It is known from a partial skeleton from the Late Triassic Wolfville Formation of Nova Scotia, described in 2003. It has many unique features seen in no other related form, including an elongated, toothless snout and large openings for the nostrils. Because of this, Teraterpeton was originally placed in its own family, Teraterpetidae, related to Trilophosaurus. Newer studies generally place it within Trilophosauridae.
Arctotraversodon is an extinct genus of traversodontid cynodonts from the Late Triassic of Canada. Fossils first described from the Wolfville Formation in Nova Scotia in 1984 represented the first known traversodontid from North America. The type and only species is A. plemmyridon and is represented by teeth and several dentary bones.
Boreogomphodon is an extinct genus of traversodontid cynodonts from the Late Triassic of the eastern United States. Fossils have been found from the Turkey Branch Formation in Virginia and the Pekin Formation of North Carolina.
Scoloparia is an extinct genus of procolophonid parareptile from the Triassic of Canada. Fossils have been found in the Early Triassic to Norian-age Wolfville Formation in Nova Scotia, Canada. Like many Triassic procolophonids, Scoloparia has expanded molar-like teeth that indicate that the animal was likely herbivorous.
Beesiiwo is a genus of hyperodapedontine rhynchosaur from the Late Triassic Popo Agie Formation in Western Wyoming. The type and only species is B. cooowuse, known from four specimens consisting of left and right maxilla fragments, and a left dentary fragment.