| Oryidae | |
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| Orya barbarica | |
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| Family: | Oryidae |
Oryidae is a monophyletic family of soil centipedes belonging to the superfamily Himantarioidea. [1]
| Oryidae | |
|---|---|
| | |
| Orya barbarica | |
| Scientific classification | |
| Kingdom: | |
| Phylum: | |
| Subphylum: | |
| Class: | |
| Order: | |
| Suborder: | |
| Superfamily: | |
| Family: | Oryidae |
Oryidae is a monophyletic family of soil centipedes belonging to the superfamily Himantarioidea. [1]
Centipedes in this family feature a short head, stout antennae, mandibles with a series of pectinate lamellae, a slightly concave labral margin with a row of denticles or bristles, uniarticulate telopodites on the first maxillae, and claws on the second maxillae fringed by two rows of filaments. The coxosternite and forcipules are short without denticles, and the forcipular segment is stout with a wide tergite. Sternal pores are mainly clustered as two pairs of groups, the ultimate legs usually have no pretarsus, and the female gonopods are distinct and usually biarticulate. [2]
These centipedes are very elongated with a high mean number of trunk segments (often greater than 100) and great variability in this number within species. [3] The number of leg-bearing segments in this family varies within as well as among species and ranges from 53 to 169. [2] The minimum number of legs recorded in this family (53 pairs) appears in the species Orphnaeus brevilabiatus ; [4] the maximum number (169 pairs) appears in the species Titanophilus maximus. [5] [2]
Centipedes in this family are found in tropical and subtropical regions of the Americas, Africa, Madagascar, south Asia, Australia, and some Pacific islands. [2]
This family contains the following genera: [6]
Pachymerium is a genus of centipedes in the family Geophilidae. Centipedes in this genus feature elongate heads, sternal pores in a pair of anterior groups and a posterior transverse band, and scattered coxal pores; the forcipular coxosternite is broad, and the ultimate article of the forcipule has a conspicuous basal denticle. These centipedes range from about 2 cm to about 8 cm in length and have 37 to 79 pairs of legs. The Chilean species Pachymerium armatum measures only 20 mm in length and has only 37 leg pairs, the minimum number observed in this genus. The Russian species P. minutum is also notable for its small size and its modest number of legs. The Portuguese species P. coiffaiti is notable for its large size and many legs. Females of the large Turkish species P. serratum can reach 78 mm in length and have as many as 79 leg pairs, the maximum number observed in this genus. This genus contains the following species:
The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae.
Himantariidae is a monophyletic family of centipedes in the order Geophilomorpha and superfamily Himantarioidea, found almost exclusively in the Northern Hemisphere. Centipedes in this family feature a short head with a concave labral margin bearing a row of denticles, a single dentate lamella and some pectinate lamellae on each mandible, second maxillae with strongly tapering telopodites and slightly spatulate claws, and a stout forcipular segment with short forcipules and a wide tergite; the ultimate legs usually have no pretarsus, and the female gonopods are distinct and biarticulate.
Mecistocephalidae is a monophyletic family of centipedes in the order Geophilomorpha. It is the only family in the suborder Placodesmata. Most species in this family live in tropical or subtropical regions, but some occur in temperate regions. This family is the third most diverse in the order Geophiliomorpha, with about 170 species, including about 130 species in the genus Mecistocephalus.
Linotaeniidae are a monophyletic clade of soil centipedes in the family Geophilidae found mostly in the temperate regions of the Holarctic as well as the south Andes. Species in the clade Linotaeniidae are characterized by a body that usually tapers toward the anterior tip; mandibles with a single pectinate lamella; second maxillae with coxo-sternite usually undivided and claws without projections; forcipular segment short, with tergite remarkably wide, forcipules evidently tapering; coxal organs opening through distinct pores on the ventral surface of the coxo-pleura. The number of legs in this clade varies within as well as among species and ranges from as few as 31 pairs of legs to as many as 83 leg pairs. Compared to most families in the suborder Adesmata, this clade features a modest number of leg-bearing segments and limited variation in this number within each species.
Schendylidae is a family of centipedes in the order Geophilomorpha.
Arrup is a genus of centipedes in the family Mecistocephalidae, native to Europe and Asia as far as Japan. Centipedes in this genus feature small telopodites on the second maxillae, the head and forcipular segment are elongate, the clypeus is almost completely areolate, and the poison glands in adult males are often deep inside the forcipules. These centipedes range from 1 cm to 5 cm in length. All species in this genus have 41 leg-bearing segments. Most are soil-dwellers but Arrup akiyoshiensis is a troglobiont.
Agnostrup is a genus of centipedes in the family Mecistocephalidae, native to Europe and Asia. Centipedes in this genus feature small telopodites without claws on the second maxillae and a robust basal denticle on the forcipular tarsungulum. These centipedes range from 2 cm to 3 cm in length. All species in this genus have 41 leg-bearing segments.
Anarrup is a genus of centipedes in the family Mecistocephalidae, native to Europe and Asia as far as Indonesia. Centipedes in this genus feature a clypeus with areolation and setae limited to a short anterior marginal band; the second maxillae have swollen and densely setose telopodites and a coxosternite divided mid-longitudinally. These centipedes range from 6 cm to 8 cm in length. All species in this genus have 41 leg-bearing segments.
Dignathodontidae is a monophyletic clade of soil centipedes in the family Geophilidae found in the Mediterranean region, extending to Macaronesia, Caucasus, and western and central Europe. The clade is characterized by a gradually anteriorly tapered body, a short head with non-attenuated antennae, and a poorly sclerotized labrum with tubercles. The number of legs in this clade varies within as well as among species and ranges from 43 pairs to 153 pairs of legs. Species in this clade tend to have more leg-bearing segments and greater intraspecific variability in this number than generally found in the family Geophilidae.
Zelanophilidae is a family of centipedes belonging to the order Geophilomorpha and superfamily Geophiloidea. Centipedes in this family are found in the Australasian region.
Gonibregmatidae are a paraphyletic family of soil centipedes belonging to the superfamily Geophiloidea.
Geophilus arenarius is a species of soil centipede in the family Geophilidae found in northwest Africa, specifically near Annaba, Algeria. This species is frequently misidentified with G. electricus, and as part of the carpophagus species-complex, is closely related to both G. carpophagus and G. easoni, though it differs mainly by lacking a transverse suture on the head and peculiar integumental features (carpophagus-structures) along the trunk, as well as having relatively stouter antennae and forcipular coxosternite. G. arenarius is distinctly smaller at full growth than G. carpophagus, with usually blunter and more sclerotised tubercles lining the intermediate part of the labrum and a minute denticle at the basis of the forcipular tarsungula. It has fewer bristles lining the lateral parts of the labrum than G. easoni as well as a generally higher number of legs and a more greyish coloured trunk. The descriptions of 36 G. arenarius syntypes indicate that males of this species have 55 pairs of legs, whereas females have 55 to 59 leg pairs, with 57 as the most common number.
Geophilus richardi is a species of soil centipede in the family Geophilidae found in France, Italy, and the Ionian Islands. Females of this species have 33 pairs of legs; males have 29 or 31. This species is one of only two in the family Geophilidae to include centipedes with as few as 29 leg pairs. This species grows up to 10 millimeters long, has no carpophagus pit or pore-fields, and has a gradually tapering, curved pretarsus of the second maxillae. G. richardi lacks typical ventral pores between 2–4mm. The sternites instead bear a small number of pores between 0.5–1mm that differ from micropores, which are unbounded by a cuticular ring. These are possibly the remnants of typical ventral pores, their smaller size being a byproduct of overall miniaturization.
Ribautia is a genus of centipedes in the family Geophilidae. It was described by French myriapodologist Henry Wilfred Brolemann in 1909. Centipedes in this genus are found in South America, tropical Africa, Madagascar, the Arabian peninsula, Australia, New Zealand, and Melanesia.
Mecistocephalus is a genus of centipedes in the family Mecistocephalidae. It was described by British entomologist George Newport in 1843.
Orphnaeus is a genus of centipedes in the family Oryidae. This genus was described by Danish entomologist Frederik Vilhelm August Meinert in 1870. Centipedes in this genus are found in tropical regions.
Maoriella is a genus of centipedes in the family Geophilidae. It was described by Austrian myriapodologist Carl Attems in 1903. Species in this genus are found in New Zealand, Australia, and Tahiti.
Gonibregmatus is a genus of centipedes in the family Gonibregmatidae. It was described by British entomologist George Newport in 1843.
Pectiniunguis is a genus of centipedes in the family Schendylidae. It was described by American naturalist Charles Harvey Bollman in 1889. Centipedes in this genus feature second maxillae with claws fringed by two rows of filaments, transversally elliptical sternal pore-fields on almost all trunk segments, two pores on each coxopleuron, and ultimate legs without claws. These centipedes range from 16 mm to 67 mm in length, have 35 to 73 pairs of legs, and are found in the tropical and subtropical regions of the Americas, Fiji, and west Africa. The African species Pectiniunguis minutus is notable not only for being the smallest in this genus but also for having as few as 35 leg pairs, the minimum number in this genus. The Brazilian species P. ducalis is notable not only for being the largest in this genus but also for having as many as 73 leg pairs, the maximum number in this genus.
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