Paramachaerodus Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Suborder: | Feliformia |
Family: | Felidae |
Subfamily: | † Machairodontinae |
Tribe: | † Smilodontini |
Genus: | † Paramachaerodus Pilgrim, 1913 |
Type species | |
Paramachaerodus orientalis (Kittl, 1887) sensu Pilgrim, 1913 | |
Other species | |
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Synonyms [1] [2] | |
Paramachaerodus
synonyms of P. orientalis
synonyms of P. maximiliani
|
Paramachaerodus is an extinct genus of saber-tooth cat of the subfamily Machairodontinae, which was endemic to Eurasia during the Middle and Late Miocene from 15 to 9 Ma. [3] A 2022 phylogenetic analysis suggested that the genus may be polyphyletic. [4]
Paramacherodus is one of the earliest known true saber-toothed cats. Many fossils were discovered in Cerro de los Batallones, a Late Miocene fossil site near Madrid, Spain. One leopard-sized species is known, Paramachaerodus orientalis from the Turolian. A second species, Paramacharodus maximiliani, has been considered a synonym of Paramachaerodus orientalis by some authors, [5] but was considered a valid species in the most recent systematic revision. [1] That revision, based on an extensive morphological analysis, also determined that the species P. ogygia exhibited less derived sabertooth features than the other Paramachaerodus species and should be assigned to a separate genus, Promegantereon . [1] [6]
The animals were about 58 centimetres (23 in) high at the shoulder, similar to a leopard, but with a more supple body. The shape of its limbs suggests that it may have been an agile climber, and could have hunted relatively large prey. [7]
In 1913, Guy Ellcock Pilgrim named the new genus Paramachaerodus for the species Machaerodus orientalis, Machaerodus schlosseri, and Felis ogygia; at the time, however, he failed to designate a type species. In 1915 he described more material that he assigned to Paramachaerodus cf. schlosseri (two hemimandibles, GSI-140 and GSI-141), though Matthew (1929) noted that both of those specimens did not closely resemble others in the genus. He rectified the lack of a type species in 1931 by designating Paramachaerodus orientalis the type species, now including Paramachaerodus schlosseri as a junior synonym. [1]
However, in the intervening span of time, Miklos Kretzoi had proposed another new genus, Pontosmilus, for the species P. orientalis (also the type species), P. schlosseri, P. ogygia, P. hungaricus, and the new species Pontosmilus indicus that he described based on GSI-141. He restricted Paramachaerodus to another new species, P. pilgrimi, that he described based on GSI-140. He also proposed the genus Proamphimachairodus for the species Machairodus maximiliani. [1]
With Pilgrim's clarification of Paramachaerodus in 1931, and utter rejection of both Pontosmilus and Proamphimachairodus-he included Machairodus orientalis, Felis ogygia, and Machairodus maximiliani as species of Paramachaerodus-Pontosmilus was rendered an invalid genus, for it possessed no valid type species, and both Pontosmilus and Proamphimachairodus were designated junior synonyms of Paramachaerodus. [1]
A major review of the genus in 2010 designated P. matthewi, P. schlosseri, P. hungaricus as junior synonyms of P. orientalis, with P. maximiliani the only other valid species in Paramachaerodus, and assigning P. agygia back to Promegantereon. It also noted that "Pontosmilus" indicus (GSI-141) was a feline, and "Pontosmilus" pilgrimi (GSI-140) a machairodontine not of Paramachaerodus or Promegantereon, though the authors offered no alternate genus assignments for either species. [1]
A third species, Paramachaerodus transasiaticus, was described in 2017 based on analysis of new fossil material from the late Miocene localities of Hezheng, Gansu Province, China, and Hadjidimovo, Bulgaria. These specimens had sabertooth characteristics intermediate between those of P. ogygia and those of P. orientalis and P. maximiliani. [8]
In 2022, a new species Paramachaerodus yingliangi was proposed based on fossils from northeastern China; the same paper also proposed separating Paramachaerodus schlosseri as the new type species and moving P. orientalis and P. maximiliani to the resurrected genus Pontosmilus. [9]
The position of Paramachaerodus within the Machairodontinae has been subject to much controversy, partially because many names have historically been proposed for it based on only scanty material. While Paramachaerodus is generally accepted as a close relative and probable forerunner of Megantereon and therefore an ancestor of Smilodon , the role Promegantereon played in this is still controversial. While researchers have generally favoured the classification of Promegantereon as a distinct genus starting in the early 2000's, it was still thought to be a close relative and potential ancestor of Paramachaerodus. Thus, Paramachaerodus' position within the Smilodontini was generally understood as intermediate, with Promegantereon as the first and Smilodon as the ultimate representative of that machairodontine tribe. [6]
However, a 2022 phylogenetic analysis by Jiangzuo et al. cast doubt on this, and indeed on the monophyly of Paramachaerodus itself. While it confirmed P. orientalis as basal to the Megantereon-Smilodon clade (the Smilodontini sensu stricto) as proposed earlier, P. maxilmiliani was instead found to be basal to a clade comprising Rhizosmilodon and Dinofelis . At the same time, Promegantereon and Paramachaerodus transasiaticus were found to group together with Metailurus and Yoshi , which are usually placed in the separate Metailurini tribe. This would make Paramachaerodus, the Smilodontini and the Metailurini as traditionally definded polyphyletic, and calls for more work regarding machairodontine systematics, suggesting that a major revision of the subfamily may be needed. [4]
Based on the morphology of its humerus, P. orientalis is inferred to have been adapted for wooded environments. [10]
Machairodus is a genus of large machairodont or ''saber-toothed cat'' that lived in Africa, Eurasia and North America during the late Miocene. It is the animal from which the subfamily Machairodontinae gets its name and has since become a wastebasket taxon over the years as many genera of sabertooth cat have been and are still occasionally lumped into it.
Dinofelis is an extinct genus of machairodontine, usually classified in the tribe Metailurini. It was widespread in Europe, Asia, Africa and North America from 5 million to about 1.2 million years ago. Fossils very similar to Dinofelis from Lothagam range back to around 8 million years ago, in the Late Miocene.
Homotherium is an extinct genus of scimitar-toothed cat belonging to the extinct subfamily Machairodontinae that inhabited North America, Eurasia, and Africa during the Pliocene and Pleistocene epochs from around 4 million to 12,000 years ago. It was one of the last surviving members of the subfamily alongside the more famous sabertooth Smilodon, to which it was distantly related. It was a large cat, comparable in size to a lion, functioning as an apex predator in the ecosystems it inhabited. In comparison to Smilodon, the canines of Homotherium were shorter, and it is suggested to have a different ecology from Smilodon as pursuit predator adapted to running down large prey in open habitats, with species of the genus also proposed to have engaged in cooperative hunting.
A saber-tooth is any member of various extinct groups of predatory therapsids, predominantly carnivoran mammals, that are characterized by long, curved saber-shaped canine teeth which protruded from the mouth when closed. Saber-toothed mammals have been found almost worldwide from the Eocene epoch to the end of the Pleistocene epoch.
Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae. They were found in Asia, Africa, North America, South America, and Europe, with the earliest species known from the Middle Miocene, with the last surviving species becoming extinct around Late Pleistocene-Holocene transition.
Metailurini is an extinct taxonomic tribe of large saber-toothed cats that lived in Africa, Asia, Europe, and North America from the Miocene to the Pleistocene.
Smilodontini is an extinct tribe within the Machairodontinae or "saber-toothed cat" subfamily of the Felidae. The tribe is also known as the "dirk-toothed cats". They were endemic to South America, North America, Europe, Asia, and Africa during the Miocene to Pleistocene, from 10.3 mya—11,000 years ago, existing for approximately 10.3 million years.
Xenosmilus is an extinct genus of homotherin machairodontine that was discovered in Florida in 2001. It has one species, Xenosmilus hodsonae.
Megantereon is an extinct genus of prehistoric machairodontine saber-toothed cat that lived in North America, Eurasia, and Africa from the late Pliocene to the Middle Pleistocene. It is a member of the tribe Smilodontini, and closely related to and possibly the ancestor of the famous American sabertooth Smilodon. In comparison to Smilodon it was somewhat smaller, around the size of a jaguar, though it is thought to have had a similar hunting strategy as an ambush predator.
Adelphailurus is an extinct genus of metailurin machairodontine (saber-toothed) cat that inhabited western North America during the middle Pliocene. It is monotypic, containing only the species Adelphailurus kansensis.
Miomachairodus is an extinct genus of large machairodontine containing only a single species, Miomachairodus pseudailuroides. It is mainly known from Middle Miocene-age fossils in Turkey and persisted until the early Late Miocene (Vallesian). Fossils of this machairodont have been found in the Vallesian-age Bahe Formation in Shaanxi, China, and Yeni Eskihisar in Anatolia. This Turkish site is of Miocene age and is well known for its pollen studies.
Lokotunjailurus is an extinct genus of saber-toothed cats (Machairodontinae) which existed during the late Miocene and earliest Pliocene epoch and is known from localities in northern, central, eastern and southern Africa. A big cat, it was more slender than comparable recent species and its build suggests cursoriality. It is grouped among a group of similar-looking saber-toothed cats known as the scimitar-tooths.
Cerro de los Batallones is a hill at Torrejón de Velasco, Madrid, Spain where a number of fossil sites from the Upper Miocene (MN10) have been found. Nine sites have been discovered with predominantly vertebrate fossils, invertebrates and plants being less represented. The first deposits were discovered accidentally in July 1991.
Homotherini (Machairodontini) is a tribe of saber-toothed cats of the family Felidae. The tribe is commonly known as scimitar-toothed cats. These saber-toothed cats were endemic to North America, Europe, Asia, Africa, and South America from the Miocene to Pleistocene living from c. 23 Ma until c. 12,000 years ago. The evolutionary relationship between the tribes Homotherini and Machairodontini cause paleontologists to classify Homotherini either as a subtribe of Machairodontini, or the same tribe often using either name interchangeably.
Rhizosmilodon is an extinct genus of saber-tooth cat of the subfamily Machairodontinae that lived during the Early Pliocene and was discovered in the U.S. state of Florida. Comparable in size to a medium-sized modern jaguar at about 165 lb. in weight, fossils of Rhizosmilodon are known only from Florida. The best specimens for this species are its lower jaw, a single skull, and teeth, which carry intermediate characteristics between advanced forms such as Smilodon and primitive forms such as Paramachairodus. It was likely an ambush predator, preying on animals such as deer, tapirs and horses.
Amphimachairodus is an extinct genus of large machairodonts. It is also a member of the tribe Homotherini within Machairodontinae and is most closely related to such species as Xenosmilus, Homotherium itself, and Nimravides. It inhabited Eurasia, Northern Africa and North America during the late Miocene epoch.
Promegantereon is an extinct genus of machairodont from the Miocene of Europe. It is one of the oldest machairodont cat species in the Smilodontini and is believed to be an ancestor of Megantereon and Smilodon.
Yoshi is an extinct genus of machairodontine sabertooth cat in the tribe Metailurini. Its fossils were described from Turolian deposits from the Miocene epoch of the Balkan Peninsula in 2014 and specimens from China once thought to belong to Metailurus. The name comes from that of the lead author's pet cat. It has been described as potentially being synonymous with Metailurus, though this is difficult to confirm at present. The type specimen is a skull that bears remarkable similarities with the modern cheetah. Yoshi is intermediate in size between a lynx and cougar, and based on several as-yet unpublished skeletons, may have had a similar lifestyle to the cheetah, being better built for speed and fast pursuit than most other machairodonts, which were more suited to ambush and hunting large, relatively slow moving animals.
Tchadailurus is a genus of machairodontine felid from the late Miocene of Chad, Africa.
Sivasmilus is a fossil genus of barbourofelid containing only a single species, Sivasmilus copei. It is known from only a single specimen, a partial mandible collected from the Chinji Formation in the Lower Siwaliks in Pakistan, estimated to be from the Miocene. The fossil was originally described in 1915 when it was assigned to the fossil feline Sivaelurus chinjiensis, but was used as the basis of a new genus and species in 1929 by Hungarian paleontologist Miklós Kretzoi. Sivasmilus copei was a relatively small, cat-like animal.