The Cupedidae are a small family of beetles, notable for the square pattern of "windows" on their elytra, which give the family their common name of reticulated beetles.
Tanyderidae, sometimes called primitive crane flies, are long, thin, delicate flies with spotted wings, superficially similar in appearance to some Tipulidae, Trichoceridae, and Ptychopteridae. Most species are restricted in distribution. They are found in many parts of the world, including North America, South America, Africa, Australia, New Zealand, and various islands in the Pacific Ocean. Adults are usually found hanging from vegetation near streams. Larvae are found either in sandy stream margins or in wet, rotten wood. Fossil species are known.
Bittacidae is a family of scorpionflies commonly called hangingflies or hanging scorpionflies.
The Ptychopteridae, phantom crane flies, are a small family of nematocerous Diptera. Superficially similar in appearance to other "tipuloid" families, they lack the ocelli of the Trichoceridae, the five-branched radial vein of the Tanyderidae, and the two anal veins that reach the wing margins of the Tipulidae. They are usually allied with the Tanyderidae based on similarities of the mesonotal suture, this group being called the Ptychopteromorpha.
Orussoidea is a superfamily of sawflies. It contains the living family Orussidae, as well as the extinct families Burmorussidae and Paroryssidae. They are the group of sawflies closest to the Apocrita, the group containing wasps, bees and ants, with both groups together forming the clade Euhymenoptera. Like most members of Apocrita, but unlike other sawflies, members of the superfamily are parasitoids.
Anaxyelidae is a family of incense cedar wood wasps in the order Hymenoptera. It contains only one living genus, Syntexis, which has only a single species, native to Western North America. Fossils of the family extend back to the Middle Jurassic, belonging to over a dozen extinct genera, with a particularly high diversity during the Early Cretaceous. Syntexis lay eggs in the sapwood of conifers, preferring recently burnt wood.
Osmylidae are a small family of winged insects of the net-winged insect order Neuroptera. The osmylids, also called lance lacewings, stream lacewings or giant lacewings, are found all over the world. There are around 225 extant species.
Pelecinidae is a family of parasitic wasps in the superfamily Proctotrupoidea. It contains only one living genus, Pelecinus, with three species known from the Americas. The earliest fossil species are known from the Jurassic, and the group was highly diverse during the Cretaceous. Members of Pelecinus are parasitic on larval beetles, flies, green lacewings, and sawflies.
Nemestrinidae, or tangle-veined flies is a family of flies in the superfamily Nemestrinoidea, closely related to Acroceridae. The family is small but distributed worldwide, with about 300 species in 34 genera. Larvae are endoparasitoids of either grasshoppers (Trichopsideinae) or scarab beetles (Hirmoneurinae). Some are considered important in the control of grasshopper populations. Adults are often observed on flowers.
The Berothidae are a family of winged insects of the order Neuroptera. They are known commonly as the beaded lacewings. The family was first named by Anton Handlirsch in 1906. The family consists of 24 genera and 110 living species distributed discontinuously worldwide, mostly in tropical and subtropical regions. Numerous extinct species have also been described. Their ecology is poorly known, but in the species where larval stages have been documented, the larvae are predators of termites.
Mesoraphidiidae is an extinct family of snakeflies in the suborder Raphidiomorpha. The family lived from the Late Jurassic through the Late Cretaceous and is known from twenty-five genera. Mesoraphidiids have been found as both compression fossils and as inclusions in amber. The family was first proposed in 1925 by the Russian paleoentomologist Andrey Vasilyevich Martynov based on Upper Jurassic fossils recovered in Kazakhstan. The family was expanded in 2002 by the synonymizing of several other proposed snakefly families. The family was divided into three subfamilies and one tribe in a 2011 paper, further clarifying the relationships of the included genera.
Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now considered to be the closest relatives of clade containing Orussoidea and Apocrita. There are about 7 genera in Karatavitidae.
Pseudopolycentropodidae is an extinct family of scorpionflies known from the Mesozoic. Fossils are known from the Middle Triassic (Anisian) to the early Late Cretaceous (Cenomanian). It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Aneuretopsychidae is an extinct family of scorpionflies known from the Mesozoic. Fossils are known from the Jurassic (Callovian-Oxfordian) to the early Late Cretaceous (Cenomanian). It is part of Mesopsychoidea, a group of scorpionflies with siphonate proboscis. They are suggested to have been nectarivores, feeding off the liquid pollination drops of and acting as pollinators for now extinct insect pollinated gymnosperms such as Bennettitales.
Mesoblattinidae is an extinct family of cockroaches known from the Mesozoic. It was formerly considered a wastebasket taxon for Mesozoic cockroaches, but the family has subsequently been better defined, with many taxa transferred to Caloblattinidae. It is considered to have close affinities with Blattidae and Ectobiidae, as well as possibly Blaberidae. The family first appeared by the Early Jurassic. They are considered to represent amongst the oldest groups of modern cockroaches, and like them are thought to have laid oothecae.
Rhagionemestriidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It was first named as a subfamily of the Nemestrinidae by Ussatchov (1968), and was raised to full family status by Nagatomi and Yang (1998). They are considered to be closely related to the family Acroceridae. Similar to Acroceridae, members of the family possess a large hemispherical head, with eyes covering nearly all of the area.
Eremochaetidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods of Asia. It is part of the extinct superfamily Archisargoidea. The morphology of the ovipositor of the only 3 dimensionally preserved genus Zhenia was initially interpreted as evidence of being an endoparasitoid of arthropods, however a subsequent study suggested that the ovipositor was used to deposit its eggs in plant material, similar to members of Tephritoidea. In a phylogenetic analysis, Ermochaetidae was found to be monophyletic, surrounded by a paraphyletic Archisargidae.
Archisargidae is an extinct family of brachyceran flies known from the Jurassic and Cretaceous periods. It is part of the extinct superfamily Archisargoidea. Most members of the family are known from the Callovian-Oxfordian Daohugou biota of Inner Mongolia, China, and the equivalently aged Karabastau Formation of Kazakhstan. The family has been found to be paraphyletic with respect to Eremochaetidae in a cladistic analysis.
Mesochrysopidae is an extinct family of lacewings known from the Jurassic and Cretaceous periods. They are considered to be closely related to green lacewings of the family Chrysopidae. The family are also alternatively considered a paraphyletic grade leading up to crown Chrysopidae.
Orthophlebiidae is an extinct family of scorpionflies known from the Triassic to Cretaceous, belonging to the superfamily Panorpoidea. The family is poorly defined and is probably paraphyletic, representing many primitive members of Panorpoidea with most species only known from isolated wings, and has such been considered a wastebasket taxon.
