Ptychoparia Temporal range: | |
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Ptychoparia striata from Czech Republic, at the National Museum (Prague) | |
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Genus: | Ptychoparia Hawle and Corda, 1847 |
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Conocephalus striatus [1] | |
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Ptychoparia is a genus of ptychopariid trilobites, and is the type genus of the family Ptychopariidae, and the order Ptychopariida.
Fossils of the various species are found in Middle Cambrian-aged marine strata of Eurasia and North America, about 513 ± 0.3 to 499 ± 1.7 million years ago.
Fossils have been found in Middle Cambrian marine strata of the Czech Republic, India, Mexico, Norway, South Korea, the United Kingdom, United States (Montana, New York, Virginia, and possibly Pennsylvania).
Several species of trilobite that were originally assigned to Ptychoparia have now been moved to other genera. [1]
Like all Ptychopariidae, the dorsal exoskeleton in Ptychoparia has an inverse egg-shaped outline. The central raised axis, that is defined by a furrow, is moderately convex and narrow, at its widest about ⅛ of the maximum width of the headshield (or cephalon). The part of the axis in the cephalon, called glabella, tapers forward, is rounded-truncate at its front, is inserted by for pairs of furrows that do not contact across the midline, and one at the back that defines the occipital ring. The fourth pair of lateral furrows curve strongly inward until parallel with the midline. A border is defined by a furrow running parallel to the margin of the cephalon, and this border furrow is deep at the front. The areas outside the axis are almost flat. Between the front of the glabella and the border, is a somewhat convex area of the cheek that is comparable in length (along the midline) to the occipital ring. In well preserved specimens the cheeks in front of the eye are adorned with very fine ridges (or striae) that probably represent parts of the digestive system (see Kordule, 2006, p. 283, figure 2.a/f). The eye ridges are threadlike, evenly curved, extending from the anterior corners of the glabella to halfway the genal angles, while the visual surface makes an abrupt angle, is shaped like a bracket parallel to the midline and ±⅛ of the length of the cephalon. Like in most trilobites, the division between the outer free cheek (or librigena) and the inner fixed cheek (or fixigena) is a line (or suture) where the exoskeleton splits to assist in moulting. It always follows the inside of the visual surface of the eye. In Ptychoparia, the part of this suture behind the eye cuts to the posterior margin just to the inside of the genal spine (or the suture is gonatoparian), and the part in front of the eye diverges from the midline. There is a spine in the back corners of the cephalon (or genal spine) that reaches approximately to the third thorax segment, and is confluent with the outer margin of the cephalon. The articulate midsection of the body (or thorax) has 13 (P. striata) or 14 (P. dubinka) segments. Tail shield (or pygidium) is slightly over half as wide as the cephalon (a state called micropygous). The pygidial axis (or rachis) has 5 or 6 rings and a termination and 4 or 5 ribs in the outlying pleural regions. The pygidium in Ptychoparia lacks a border. [2]
Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.
Phacopidae is a family of phacophid trilobites that ranges from the Lower Ordovician to the Upper Devonian, with representatives in all paleocontinents.
Paradoxides is a genus of large to very large trilobite found throughout the world during the Middle Cambrian period. One record-breaking specimen of Paradoxides davidis, described by John William Salter in 1863, is 37 cm (15 in). The cephalon was semicircular with free cheeks ending in long, narrow, recurved spines. Eyes were crescent shaped providing an almost 360° view, but only in the horizontal plane. Its elongate thorax was composed of 19-21 segments and adorned with longish, recurved pleural spines. Its pygidium was comparatively small. Paradoxides is a characteristic Middle-Cambrian trilobite of the 'Atlantic' (Avalonian) fauna. Avalonian rocks were deposited near a small continent called Avalonia in the Paleozoic Iapetus Ocean. Avalonian beds are now in a narrow strip along the East Coast of North America, and in Europe.
Triarthrus is a genus of Upper Ordovician ptychopariid trilobite found in New York, Ohio, Kentucky, and Indiana, eastern and northern Canada, China and Scandinavia. It is the last of the Olenid trilobites, a group which flourished in the Cambrian period. The specimens of T. eatoni that are found in the Beecher's Trilobite Bed, Rome, New York area are exquisitely preserved showing soft body parts in iron pyrite. Pyrite preservation has given scientists a rare opportunity to examine the gills, walking legs, antennae and digestive systems of trilobites, which are rarely preserved. Triarthrus is therefore commonly used in science texts to illustrate trilobite anatomy and physiology.
Asaphus is a genus of trilobites that is known from the Lower and Middle Ordovician of northwestern Europe.
Dikelocephalus is a genus of very large trilobites of up to 50 cm (20 in) long, that lived during the last 3 million years of the Cambrian (Sunwaptan). Their fossils are commonly found as disarticulated sclerites, in the upper Mississippi Valley and in Canada (Alberta). The exoskeleton is rounded anteriorly, with the thorax and sides of the tailshield slightly tapering to about ⅔× of the width across the base of the spines at the back of the headshield. At the side corners of the pygidium there may be triangular or hooked spines, pointing backwards, while between the spines the posterior margin is at a 30-75° angle with the lateral margin, gently convex or nearly straight. If pygidial spines are lacking, the margin is gradually rounded. The thorax has 12 segments.
Acimetopus Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi (1943), Order Agnostida Salter (1864). It lived during the Botomian stage. = late Lower Cambrian Stage 4 ; the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage.
Meniscuchus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the Botomian stage, which lasted from approximately 522 to 516 million years ago. This faunal stage was part of the Cambrian Period. Meniscuchus has been found in the USA, Canada, Russia and Australia.
Dicerodiscus is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the early part of the Botomian stage, in China. Four species have been assigned to it. Dicerodiscus is unique for an eodiscoid in having conspicuous and curved spines that are attached anteriorly, and at their base are directed outward perpendicular to the midline, before gradually bending further backwards.
Yunnanocephalus is a genus of ptychopariid trilobite. It lived during the late Atdabanian and Botomian stages, in what are currently Antarctica, Australia and China. It was a "moderately common" member of the Chengjiang Fauna. Yunnanocephalus is the only genus currently assigned to the Yunnanocephalidae family.
Meteoraspis is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae. The various species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Meteoraspis are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as M. nevensis from Victoria Land, Antarctica.
Eoredlichia is an extinct genus of trilobite of average to large size. It lived during the early Cambrian in the Chengjiang fauna of Yunnan, China, and in Australia and Thailand. Eoredlichia is compounded of the Greek ἠώς and Redlichia, a later but related genus, so it means "early Redlichia". The species epithet intermedia means intermediate, indicating it is morphologically intermediate between other species. Eofallotaspis gives rise to Lemdadella, and thence to Eoredlichia and the other Redlichiidae.
Holmiidae is a family of trilobites, that lived during the Lower Cambrian (Atdabanian). The Holmiidae is a diverse family of eight genera containing at least 17 species. It includes some of the earliest trilobites of Baltica. Holmiidae occur throughout Baltica and Western Laurentia, and also in Morocco.
Alokistocaridae is a family of ptychopariid trilobites that lived from the Botomian epoch of the Early Cambrian until the Late Cambrian. Alokistocarids were particle feeders and left small furrows which are occasionally preserved. Their remains are found worldwide. Elrathia kingii, one of the most collected trilobites in the world, is a typical alokistocarid.
Conocoryphe is a genus of primarily eyeless trilobites belonging to the family Conocoryphidae. They lived during the Middle Cambrian period, about 505 million years ago. These arthropods lived on the sea bottom (epifaunal) and lived off dead particulate organic matter.
Ovatoryctocara is a genus of small corynexochid trilobites from the Cambrian, that lived in what now are Siberia, China, Greenland and Canada (Newfoundland). Ovatoryctocara can be recognised by the combination of the following characters: the central raised area of the cephalon is approximately cylindrical and has two rows of four triangular or round pits. The thorax only has 5 or 6 segments. The tailshield has an axis of 6 to 12 rings, the pleural furrows are well developed and the border is absent or narrow as a hair.
Cedaria is a small, rather flat trilobite with an oval outline, a headshield and tailshield of approximately the same size, 7 articulating segments in the middle part of the body and spines at the back edges of the headshield that reach halflength of the body. Cedaria lived during the early part of the Upper Cambrian (Dresbachian), and is especially abundant in the Weeks Formation.
Tricrepicephalus is an extinct genus of ptychopariid trilobites of the family Tricrepicephalidae with species of average size. Its species lived from 501 to 490 million years ago during the Dresbachian faunal stage of the late Cambrian Period. Fossils of Tricrepicephalus are widespread in Late Cambrian deposits in North America, but is also known from one location in South-America. Tricrepicephalus has an inverted egg-shaped exoskeleton, with three characteristic pits in the fold that parallels the margin of the headshield just in front of the central raised area. The articulating middle part of the body has 12 segments and the tailshield carries two long, tubular, curved pygidial spines that are reminiscent of earwig's pincers that rise backwards from the plain of the body at approximately 30°.
Anabaraspis is a genus of redlichiid trilobite. A. splendens occurs in the uppermost Lower Cambrian and lowest Middle Cambrian of Russia. In Anabaraspis there is a long area in front of the glabella which is not differentiated in a border and a preglabellar field. This is a unique character in the family Paradoxididae. The frontal lobe of the central raised area of the headshield is slightly pointed, rather than rounded or truncate, a character shared with Plutonides, though, in Plutonides it hangs over the short anterior border.
Viaphacops is a genus of trilobites in the order Phacopida, family Phacopidae, that lived during the Middle Devonian, and is known from North and South America, Asia.