Stomiati | |
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Illustration of a stoplight loosejaw (M. niger) | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Clade: | Euteleostei |
Clade: | Stomiati |
Orders | |
Stomiati is a group of teleost fish belonging to the cohort (group) Euteleostei , which is a group of bony fishes within the infra-class Teleostei that evolved ~240 million years ago. Teleostei is a group of ray-finned fishes with the exception of primitive bichirs, sturgeons, paddlefishes, freshwater garfishes, and bowfins. The cohort of Euteleostei is divided into two smaller groups: the Protacanthopterygii and the Neoteleostei . Stomiati happen to be descendants of the Protacanthopterygii, and contains the order of Osmeriformes and Stomiiformes (Betancur-R, et al.). [1]
Stomiati is one of five major euteleost lineages that were recently placed in one monophyly, a group of organisms all descending from one common ancestor. The five being a clade formed by Esociformes and Salmoniformes; second being the Stomiatii consisting of only Osmeriformes and Stomiiformes; Argentiniformes (excludes Alepocephaliformes); Galaxiiformes (excludes Lepidogalaxias ); and Neotelestei. Stomiati are known for their large mouth, long throat/barbel, and no caudal filament. [2]
Osmeriformes smelts belong to the subcohort Protacanthopterygii. Osmeriformes mainly spawn in freshwater habitats except for the osmerid (family Osmeridae) Osmerus eperlanus , and one or two salangids (family Salangidae). Fourteen different families comprise the order of Osmeriformes. [3]
Stomiiformes are an order of deep sea ray-finned fishes. These fishes are known, and named, for their absurdly large mouth openings. Stomiiformes is derived from Ancient Greek meaning “mouth” + “external form” which references their large mouth openings. Four (some may claim 5) different families comprise the order of Stomiiformes. [4]
The order of stomiiformes is linked to some of the most interesting deep sea dragonfishes. The genera of Malacosteus , Aristostomias and Pachystomias are the only fishes that have the ability to produce red bioluminescence. The species Malacosteusniger is found lurking below a depth of 500 m (1,600 ft) illuminating its red light in order pick out prey. The reason this red bioluminescent light is so interesting is the fact that below the depth of 500 meters, there are very few organisms that have the ability to perceive this wavelength. Therefore, M. niger effortlessly hunts for prey with the aid of light while its prey remains in seemingly complete darkness.
Little is known about their ability to use and let alone sense light at such depths. Due to them lurking at a depth well below 500 m, they can only be observed in their natural habitat, and bringing one up to the surface alive is infeasible. The change in pressure the fish would experience would be too great and most certainly be fatal. [5]
Actinopterygii, members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species.
Osteichthyes, commonly referred to as the bony fish, is a diverse superclass of vertebrates, that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilage. The vast majority of fish are members of Osteichthyes, which is an extremely diverse and abundant group consisting of 45 orders, and over 435 families and 28,000 species. It is the largest class of vertebrates in existence today. The group Osteichthyes is divided into the ray-finned fish (Actinopterygii) and lobe-finned fish (Sarcopterygii). The oldest known fossils of bony fish are about 425 million years old, which are also transitional fossils, showing a tooth pattern that is in between the tooth rows of sharks and bony fishes.
Deep-sea fish are fish that live in the darkness below the sunlit surface waters, that is below the epipelagic or photic zone of the sea. The lanternfish is, by far, the most common deep-sea fish. Other deep sea fishes include the flashlight fish, cookiecutter shark, bristlemouths, anglerfish, viperfish, and some species of eelpout.
The jellynose fishes or tadpole fishes are the small order Ateleopodiformes. This group of ray-finned fish is monotypic, containing a single family Ateleopodidae. It has about a dozen species in four genera, but these enigmatic fishes are in need of taxonomic revision.
Osteoglossiformes is a relatively primitive order of ray-finned fish that contains two sub-orders, the Osteoglossoidei and the Notopteroidei. All of at least 245 living species inhabit freshwater. They are found in South America, Africa, Australia and southern Asia, having first evolved in Gondwana before that continent broke up. In 2008 several new species of marine osteoglossiforms was described from the Danish Eocene Fur Formation dramatically increases the diversity of this group. This implies that the Osteoglossomorpha is not a primary freshwater fish group with the osteoglossiforms having a typical Gondwana distribution.
Neopterygii is a subclass of ray-finned fish (Actinopterygii). Neopterygii includes the Holostei and the Teleostei, of which the latter comprise the vast majority of extant fishes, and over half of all living vertebrate species. While living holosteans include only freshwater taxa, teleosts are diverse in both freshwater and marine environments. Many new species of teleosts are scientifically described each year.
The Osmeriformes are an order of ray-finned fish that includes the true or freshwater smelts and allies, such as the galaxiids and noodlefishes; they are also collectively called osmeriforms. They belong to the teleost superorder Protacanthopterygii, which also includes pike and salmon, among others. The order's name means "smelt-shaped", from Osmerus + the standard fish order suffix "-formes". It ultimately derives from Ancient Greek osmé + Latin forma, the former in reference to the characteristic aroma of the flesh of Osmerus.
Stomiiformes is an order of deep-sea ray-finned fishes of very diverse morphology. It includes, for example, dragonfishes, lightfishes, loosejaws, marine hatchetfishes and viperfishes. The order contains 4 families with more than 50 genera and at least 410 species. As usual for deep-sea fishes, there are few common names for species of the order, but the Stomiiformes as a whole are often called dragonfishes and allies or simply stomiiforms.
The Myctophiformes are an order of ray-finned fishes consisting of two families of deep-sea marine fish, most notably the highly abundant lanternfishes (Myctophidae). The blackchins (Neoscopelidae) contain six species in three genera, while the bulk of the family belongs to the Myctophidae, with over 30 genera and some 252 species.
The marine hatchetfishes or deep-sea hatchetfishes as well as the related bottlelights, pearlsides and constellationfishes are small deep-sea ray-finned fish of the stomiiform family Sternoptychidae. They are not closely related to and should not be confused with the freshwater hatchetfishes, which are teleosts in the characiform family Gasteropelecidae. The Sternoptychidae have 10 genera and about 70 species altogether.
Lepidogalaxias salamandroides is a species of small fish of Western Australia. It is the only member of the family Lepidogalaxiidae and genus Lepidogalaxias. Common names for this fish include salamanderfish and Shannon mudminnow. Although it is not a lungfish, it resembles lungfish in several respects, including its ability to survive dry seasons by burrowing into the sand. It is on the IUCN Red List as Endangered.
A viperfish is any species of marine fish in the genus Chauliodus. Viperfishes are mostly found in the mesopelagic zone and are characterized by long, needle-like teeth and hinged lower jaws. A typical viperfish grows to lengths of 30 cm (12 in). Viperfishes undergo diel vertical migration and are found all around the world in tropical and temperate oceans. Viperfishes possess photophores along the ventral side of their body, likely used to camouflage them by blending in with the less than 1% of light that reaches to below 200 meters depth.
Stomiidae is a family of deep-sea ray-finned fish, including the barbeled dragonfishes. They are quite small, usually around 15 cm, up to 26 cm. These fish are apex predators and have enormous jaws filled with fang-like teeth. They are also able to hinge the neurocranium and upper-jaw system, which leads to the opening of the jaw to more than 100 degrees. This ability allows them to consume extremely large prey, often 50% greater than their standard length.
The stoplight loosejaws are small, deep-sea dragonfishes of the genus Malacosteus, classified either within the subfamily Malacosteinae of the family Stomiidae, or in the separate family Malacosteidae. They are found worldwide, outside of the Arctic and Subantarctic, in the mesopelagic zone below a depth of 500 meters. This genus once contained three nominal species: M. niger, M. choristodactylus, and M. danae, with the validity of the latter two species being challenged by different authors at various times. In 2007, Kenaley examined over 450 stoplight loosejaw specimens and revised the genus to contain two species, M. niger and the new M. australis.
Malacosteus niger, commonly known as the black dragon fish, is a species of deep-sea fish. Some additional common names for this species include: northern stoplight loosejaw, lightless loosejaw, black loosejaw, and black hinged-head. It belongs to the family Stomiidae, or dragonfishes. It is among the top predators of the open mesopelagic zone. M. niger is a circumglobal species, which means that it inhabits waters ranging from the tropics to the subarctics. Not many studies have been conducted on its feeding habits, but recent research suggests that M. niger primarily feed on calanoid copepods which is a form of zooplankton. Indeed, it appears that M. niger primarily prey on zooplankton despite its apparent morphological adaptations for the consumption of relatively large prey. Another unique adaptation for this species is its ability to produce both red and blue bioluminescence. Most mesopelagic species aren't capable of producing red bioluminescence. This is advantageous because most other species cannot perceive red light, therefore allowing M. niger to camouflage part of itself to its prey and predators.
Protacanthopterygii is a ray-finned fish taxon ranked as a superorder of the infraclass Teleostei. They inhabit both marine and freshwater habitats. They appear to have evolved in the Cretaceous or perhaps late Jurassic, originating probably roughly 150 million years ago; fossils of them and the closely related Otocephala are known from throughout the Cretaceous.
The Argentiniformes are an order of ray-finned fish whose distinctness was recognized only fairly recently. In former times, they were included in the Osmeriformes as suborder Argentinoidei. That term refers only to the suborder of marine smelts and barreleyes in the classification used here, with the slickheads and allies being the Alepocephaloidei. These suborders were treated as superfamilies Argentinoidea and Alepocephaloidea, respectively, when the present group was still included in the Osmeriformes.
Aristostomias is a genus of barbeled dragonfishes native to the ocean depths in the Pacific, Atlantic and Indian oceans.
Pachystomias microdon, the smalltooth dragonfish, is a species of barbeled dragonfish found in the oceans at depths of from 660 to 4,000 metres. This species grows to a length of 22.1 centimetres (8.7 in) SL. This species is the only known species in its genus.
Otocephala is a clade of ray-finned fishes within the infraclass Teleostei that evolved some 230 million years ago. It is named for the presence of a hearing (otophysic) link from the swimbladder to the inner ear. Other names proposed for the group include Ostarioclupeomorpha and Otomorpha.