Thyreosaurus Temporal range: Middle Jurassic | |
---|---|
Skeletal reconstruction without dorsal plates | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Clade: | Dinosauria |
Clade: | † Ornithischia |
Clade: | † Thyreophora |
Clade: | † Stegosauria |
Family: | † Stegosauridae |
Subfamily: | † Dacentrurinae |
Genus: | † Thyreosaurus Zafaty et al., 2024 |
Species: | †T. atlasicus |
Binomial name | |
†Thyreosaurus atlasicus Zafaty et al., 2024 | |
Thyreosaurus is an extinct genus of stegosaurian dinosaur from the Middle Jurassic El Mers III Formation of Morocco. The genus contains a single species, Thyreosaurus atlasicus, known form a partial disarticulated partial skeleton consisting of several dorsal vertebrae, ribs, a limb bone, and dermal bones. Thyreosaurus is characterized by a unique dermal armor with an asymmetrical texture, indicating that it may have been recumbent. Alongside Adratiklit , Thyreosaurus increases the diversity of unique stegosaurs occurring in the Middle Jurassic of Africa.
The Thyreosaurus holotype specimen, HIIUC-BN00, was discovered in 2021 in sediments of the El Mers III Formation (El Mers Group, Boulahfa North locality) near Boulemane in the Middle Atlas mountains of Morocco. The specimen—which is associated but disarticulated—consists of nine dorsal vertebrae, 21 fragmentary ribs, a limb bone tentatively described as a fibula, and six osteoderms. [1]
The fossil material was first reported in several conference abstracts in 2022 and 2023 before its formal description. [2] [3] [4]
In 2024, Zafaty et al. described Thyreosaurus atlasicus as a new genus and species of stegosaurian dinosaur based on these fossil remains. The generic name, Thyreosaurus, combines thyreos (θυρεός), the name of an oblong shield used in Ancient Greece—referencing the specimen's unusual osteoderms—with the Greek σαῦρος (sauros), meaning "lizard". The specific name, atlasicus, refers to the Atlas Mountains of Morocco. [1]
The holotype of Thyreosaurus belonged to an adult individual with an estimated body length of 6 metres (20 ft), but it was not fully grown. It possessed unique dermal armor that has been compared to that of nodosaurid ankylosaurs, [4] consisting of asymmetrically sided oblong plates that were likely arranged recumbent on the back of the animal, instead of being held erect as in other stegosaurs. [1]
In their phylogenetic analysis, Zafaty et al. (2024) recovered Thyreosaurus as dacentrurine member of the Stegosauridae, as the sister taxon to Dacentrurus . These species are in turn sister to Adratiklit , a contemporary of Thyreosaurus. Their results are displayed in the cladogram below: [1]
Thyreophora is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous.
Kentrosaurus is a genus of stegosaurid dinosaur from the Late Jurassic in Lindi Region of Tanzania. The type species is K. aethiopicus, named and described by German palaeontologist Edwin Hennig in 1915. Often thought to be a "primitive" member of the Stegosauria, several recent cladistic analyses find it as more derived than many other stegosaurs, and a close relative of Stegosaurus from the North American Morrison Formation within the Stegosauridae.
Polacanthus, deriving its name from the Ancient Greek polys-/πολύς- "many" and akantha/ἄκανθα "thorn" or "prickle", is an early armoured, spiked, plant-eating ankylosaurian dinosaur from the early Cretaceous period of England.
Scelidosaurus is a genus of herbivorous armoured ornithischian dinosaur from the Jurassic of England.
Wuerhosaurus is a genus of stegosaurid dinosaur from the Early Cretaceous Period of China and Mongolia. As such, it was one of the last genera of stegosaurians known to have existed.
Huayangosaurus is a genus of stegosaurian dinosaur from the Middle Jurassic of China. The name derives from "Huayang" (華陽), an alternate name for Sichuan, and "saurus", meaning "lizard". It lived during the Bathonian to Callovian stages, around 165 million years ago, some 20 million years before its famous relative, Stegosaurus appeared in North America. At only approximately 4 metres (13 ft) long, it was also much smaller than its famous cousin. Found in the Lower Shaximiao Formation, Huayangosaurus shared the local Middle Jurassic landscape with the sauropods Shunosaurus, Datousaurus, Omeisaurus and Protognathosaurus, the ornithopod Xiaosaurus and the carnivorous Gasosaurus.
Lexovisaurus is a genus of stegosaur from mid-to-Late Jurassic Europe, 165.7-164.7 mya. Fossils of limb bones and armor fragments have been found in middle to late Jurassic-aged strata of England and France.
Stegosauria is a group of herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere, Africa and possibly South America. Their geographical origins are unclear; the earliest unequivocal stegosaurian, Bashanosaurus primitivus, was found in the Bathonian Shaximiao Formation of China.
Dracopelta is a monospecific genus of ankylosaur dinosaur from Portugal that lived during the Late Jurassic in what is now the Lourinhã Formation. The type and only species is Dracopelta zbyszewskii, which is represented by a partial skeleton including unpublished material.
Emausaurus is a genus of thyreophoran or armored dinosaur from the Early Jurassic. Its fossils have been found in Mecklenburg-Vorpommern, northern Germany. Emausaurus is the only known Toarcian thyreophoran, as well as the only dinosaur from the zone of the same age with a formal name.
Isaberrysaura is a genus of stegosaurian ornithischian dinosaur from the Middle Jurassic Los Molles Formation of Patagonia, Argentina. The genus contains a single species, I. mollensis, described by Salgado et al. in 2017 from a single specimen. Although initially classified as a basal neornithischian, subsequent analysis has allied it with the Stegosauria; the morphology of its skull resembles those of other members of the group.
The El Mers Group is a geological group in the Middle Atlas of Morocco. It is subdivided into three formations named the El Mers I, II, and III Formations, respectively. It is a marine deposit primarily consisting of marl, with gypsum present in the upper part of unit 3. It is the lateral equivalent of the terrestrial Guettioua Sandstone. Dinosaur remains are among the fossils that have been recovered from the group, most notably those of sauropods and the unusual thyreophorans Adratiklit, Thyreosaurus, and Spicomellus.
Adratiklit is an extinct genus of herbivorous stegosaurian dinosaur that lived on the supercontinent Gondwana during the Middle Jurassic period. The genus contains a single species, Adratiklit boulahfa. Its remains were found in the El Mers III Formation (Bathonian–?Callovian), near Boulahfa, south of Boulemane, Fès-Meknès, north Morocco.
Spicomellus is an extinct genus of early ankylosaurian dinosaur from the El Mers III Formation (Bathonian-Callovian) of Morocco. The genus contains a single species, S. afer, known from a single rib with fused osteoderms. Spicomellus represents the oldest named ankylosaur.
Parankylosauria is a group of basal ankylosaurian dinosaurs known from the Cretaceous of South America, Antarctica, and Australia. It is thought the group split from other ankylosaurs during the mid-Jurassic period, despite this being unpreserved in the fossil record.
Bashanosaurus is an extinct genus of stegosaurian dinosaur from the Middle Jurassic Shaximiao Formation of Yunyang County, China. The genus contains a single species, Bashanosaurus primitivus, known from incomplete skeletons belonging to three individuals. It is one of the basalmost stegosaurs, as well as one of the oldest known stegosaurs, along with Adratiklit, Isaberrysaura, and Thyreosaurus.
Jakapil is a genus of basal thyreophoran dinosaur from the Candeleros Formation of Argentina. The type species is Jakapil kaniukura.
Baiyinosaurus is an extinct genus of stegosaurian dinosaurs from the Middle Jurassic Wangjiashan Formation of China. The genus contains a single species, B. baojiensis, known from a partial skeleton including cranial bones. The skeletal anatomy of Baiyinosaurus demonstrates transitional features between basal thyreophorans and stegosaurs. While many stegosaurs are known from China, Baiyinosaurus is the only one currently named from Gansu Province.