Thyreosaurus

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Thyreosaurus
Temporal range: Middle Jurassic
(Bathonian–?Callovian)
Thyreosaurus Skeletal.svg
Skeletal reconstruction without dorsal plates
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Ornithischia
Clade: Thyreophora
Clade: Stegosauria
Family: Stegosauridae
Subfamily: Dacentrurinae
Genus: Thyreosaurus
Zafaty et al., 2024
Species:
T. atlasicus
Binomial name
Thyreosaurus atlasicus
Zafaty et al., 2024

Thyreosaurus is an extinct genus of stegosaurian dinosaur from the Middle Jurassic El Mers III Formation of Morocco. The genus contains a single species, Thyreosaurus atlasicus, known form a partial disarticulated partial skeleton consisting of several dorsal vertebrae, ribs, a limb bone, and dermal bones. Thyreosaurus is characterized by a unique dermal armor with an asymmetrical texture, indicating that it may have been recumbent. Alongside Adratiklit , Thyreosaurus increases the diversity of unique stegosaurs occurring in the Middle Jurassic of Africa.

Contents

Discovery and naming

Morocco location map.svg
Red pog.svg
Thyreosaurus type locality near Boulemane, Morocco

The Thyreosaurus holotype specimen, HIIUC-BN00, was discovered in 2021 in sediments of the El Mers III Formation (El Mers Group, Boulahfa North locality) near Boulemane in the Middle Atlas mountains of Morocco. The specimen—which is associated but disarticulated—consists of nine dorsal vertebrae, 21 fragmentary ribs, a limb bone tentatively described as a fibula, and six osteoderms. [1]

The fossil material was first reported in several conference abstracts in 2022 and 2023 before its formal description. [2] [3] [4]

In 2024, Zafaty et al. described Thyreosaurus atlasicus as a new genus and species of stegosaurian dinosaur based on these fossil remains. The generic name, Thyreosaurus, combines thyreos (θυρεός), the name of an oblong shield used in Ancient Greece—referencing the specimen's unusual osteoderms—with the Greek σαῦρος (sauros), meaning "lizard". The specific name, atlasicus, refers to the Atlas Mountains of Morocco. [1]

Description

The holotype of Thyreosaurus belonged to an adult individual with an estimated body length of 6 metres (20 ft), but it was not fully grown. It possessed unique dermal armor that has been compared to that of nodosaurid ankylosaurs, [4] consisting of asymmetrically sided oblong plates that were likely arranged recumbent on the back of the animal, instead of being held erect as in other stegosaurs. [1]

Classification

In their phylogenetic analysis, Zafaty et al. (2024) recovered Thyreosaurus as dacentrurine member of the Stegosauridae, as the sister taxon to Dacentrurus . These species are in turn sister to Adratiklit , a contemporary of Thyreosaurus. Their results are displayed in the cladogram below: [1]

Stegosauria

Related Research Articles

<i>Stegosaurus</i> Thyreophoran stegosaurid dinosaur genus from Late Jurassic period

Stegosaurus is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been found in the western United States and in Portugal, where they are found in Kimmeridgian- to Tithonian-aged strata, dating to between 155 and 145 million years ago. Of the species that have been classified in the upper Morrison Formation of the western US, only three are universally recognized: S. stenops, S. ungulatus and S. sulcatus. The remains of over 80 individual animals of this genus have been found. Stegosaurus would have lived alongside dinosaurs such as Apatosaurus, Diplodocus, Camarasaurus and Allosaurus, the latter of which may have preyed on it.

<span class="mw-page-title-main">Thyreophora</span> Extinct clade of dinosaurs

Thyreophora is a group of armored ornithischian dinosaurs that lived from the Early Jurassic until the end of the Cretaceous.

<span class="mw-page-title-main">Ankylosauria</span> Extinct order of dinosaurs

Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in the Middle Jurassic, and persisted until the end of the Cretaceous Period. The two main families of Ankylosaurs, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous.

<i>Kentrosaurus</i> Extinct genus of dinosaurs from late Jurassic in Lindi Region, Tanzania

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<i>Polacanthus</i> Extinct genus of reptiles

Polacanthus, deriving its name from the Ancient Greek polys-/πολύς- "many" and akantha/ἄκανθα "thorn" or "prickle", is an early armoured, spiked, plant-eating ankylosaurian dinosaur from the early Cretaceous period of England.

<i>Scelidosaurus</i> Genus of reptiles (fossil)

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<i>Wuerhosaurus</i> Extinct genus of dinosaurs

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<i>Lexovisaurus</i> Extinct genus of reptiles

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<span class="mw-page-title-main">Stegosauria</span> Extinct clade of dinosaurs

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<i>Emausaurus</i> Extinct genus of thyreophoran dinosaurs

Emausaurus is a genus of thyreophoran or armored dinosaur from the Early Jurassic. Its fossils have been found in Mecklenburg-Vorpommern, northern Germany. Emausaurus is the only known Toarcian thyreophoran, as well as the only dinosaur from the zone of the same age with a formal name.

The El Mers Group is a geological group in the Middle Atlas of Morocco. It is subdivided into three formations named the El Mers I, II, and III Formations, respectively. It is a marine deposit primarily consisting of marl, with gypsum present in the upper part of unit 3. It is the lateral equivalent of the terrestrial Guettioua Sandstone. Dinosaur remains are among the fossils that have been recovered from the group, most notably those of sauropods and the unusual thyreophorans Adratiklit, Thyreosaurus, and Spicomellus.

<i>Adratiklit</i> Extinct genus of dinosaurs

Adratiklit is an extinct genus of herbivorous stegosaurian dinosaur that lived on the supercontinent Gondwana during the Middle Jurassic period. The genus contains a single species, Adratiklit boulahfa. Its remains were found in the El Mers III Formation (Bathonian–?Callovian), near Boulahfa, south of Boulemane, Fès-Meknès, north Morocco.

<i>Spicomellus</i> Extinct genus of dinosaurs

Spicomellus is an extinct genus of early ankylosaurian dinosaur from the El Mers III Formation (Bathonian-Callovian) of Morocco. The genus contains a single species, S. afer, known from a single rib with fused osteoderms. Spicomellus represents the oldest named ankylosaur.

<span class="mw-page-title-main">Parankylosauria</span> Extinct group of dinosaurs

Parankylosauria is a group of basal ankylosaurian dinosaurs known from the Cretaceous of South America, Antarctica, and Australia. It is thought the group split from other ankylosaurs during the mid-Jurassic period, despite this being unpreserved in the fossil record.

<i>Bashanosaurus</i> Extinct genus of stegosaurian dinosaurs

Bashanosaurus is an extinct genus of stegosaurian dinosaur from the Middle Jurassic Shaximiao Formation of Yunyang County, China. The genus contains a single species, Bashanosaurus primitivus, known from incomplete skeletons belonging to three individuals. It is one of the basalmost stegosaurs, as well as one of the oldest known stegosaurs, along with Adratiklit, Isaberrysaura, and Thyreosaurus.

<i>Jakapil</i> Extinct genus of dinosaurs

Jakapil is a genus of basal thyreophoran dinosaur from the Candeleros Formation of Argentina. The type species is Jakapil kaniukura.

<i>Baiyinosaurus</i> Genus of stegosaurian dinosaurs

Baiyinosaurus is an extinct genus of stegosaurian dinosaurs from the Middle Jurassic Wangjiashan Formation of China. The genus contains a single species, B. baojiensis, known from a partial skeleton including cranial bones. The skeletal anatomy of Baiyinosaurus demonstrates transitional features between basal thyreophorans and stegosaurs. While many stegosaurs are known from China, Baiyinosaurus is the only one currently named from Gansu Province.

References

  1. 1 2 3 4 Zafaty, O.; Oukassou, M.; Riguetti, F.; Company, J.; Bendrioua, S.; Tabuce, R.; Charrière, A.; Pereda-Suberbiola, X. (2024). "A new stegosaurian dinosaur (Ornithischia: Thyreophora) with a remarkable dermal armour from the Middle Jurassic of North Africa". Gondwana Research . 131: 344–362. Bibcode:2024GondR.131..344Z. doi:10.1016/j.gr.2024.03.009.
  2. Zafaty, O.; Oukassou, M.; Pereda-Suberbiola, X.; Riguetti, F.; Company, J.; Bendrioua, S.; Tabuce, R.; A. Charrière (2022). New remains of thyreophoran dinosaurs (Ornithischia) from the Middle Jurassic red beds of Morocco. 4èmes Journées Jeunes Chercheurs en Géosciences.
  3. Zafaty, O.; Oukassou, M.; Pereda-Suberbiola, X.; Riguetti, F.; Company, J.; Bendrioua, S.; Tabuce, R.; A. Charrière (2023). Nouvelle découverte d'un thyréophore à armure dermique remarquable dans le Jurassique moyen de l'Afrique du Nord (Maroc). Huitième Rencontre internationale sur la valorisation et la préservation du patrimoine paléontologique. pp. 26–27.
  4. 1 2 Zafaty, Omar; Oukassou, M.; Pereda-Suberbiola, X.; Riguetti, F.; Company, J.; Bendrioua, S.; Tabuce, R.; A. Charrière (2023). A new partial skeleton including dermal armor of a thyreophoran (Ornithischia) dinosaur from Middle Jurassic of North Africa (Morocco). 14th Symposium on Mesozoic Terrestrial Ecosystems and Biota. Vol. 306. pp. 261–264. doi: 10.1002/ar.25219 .