Trogulus | |
---|---|
Trogulus tricarinatus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Chelicerata |
Class: | Arachnida |
Order: | Opiliones |
Family: | Trogulidae |
Genus: | Trogulus Latreille, 1802 |
Trogulus is a genus of Opiliones (also known as harvestmen) in the family Trogulidae. Harvestmen in the genus have large, elongated and flattened bodies (prosoma and opisthosoma) and a two-segmented tarsus segment on leg II, which distinguishes them from other genera in the family Trogulidae. The legs tend to be short compared to most harvestmen. [1] External morphology tends to be very uniform across species in the genus, making differentiation of species difficult. [2] Species occur in a range of habitats, but are most common in forests where their primary prey, snails, are abundant. Trogulus contains the largest known harvestman by body length, Trogulus torosus. [2]
The genus Trogulus is divided into seven species groups [1] [3] –groupings of allopatric species with morphological similarities: [4]
However, genetic analysis indicates that two of the seven species groups listed above are not monophyletic. The T. torosus species group with the monophyletic T. hirtus species group nested within it and the T. gypseus species group which contains the monophyletic T. tricarinatus species group are both paraphyletic.
Trogulus contains the following species: [5]
The Opiliones are an order of arachnids colloquially known as harvestmen, harvesters, harvest spiders, or daddy longlegs. As of April 2017, over 6,650 species of harvestmen have been discovered worldwide, although the total number of extant species may exceed 10,000. The order Opiliones includes five suborders: Cyphophthalmi, Eupnoi, Dyspnoi, Laniatores, and Tetrophthalmi, which were named in 2014.
Opilio is a genus of harvestmen with 35 known species.
Dyspnoi is a suborder of harvestmen, currently comprising 43 extant genera and 356 extant species, although more species are expected to be described in the future. The eight families are currently grouped into three superfamilies: the Acropsopilionoidea, Ischyropsalidioidea, and Troguloidea.
The Phalangiidae are a family of harvestmen with about 380 known species. The best known is Phalangium opilio. Dicranopalpus ramosus is an invasive species in Europe.
Leiobunum is a genus of the harvestman family Sclerosomatidae with more than 100 described species. Contrary to popular belief, they are not spiders, although they share a resemblance. They are arachnids, in the order Opiliones, harvestmen. Species in Leiobunum tend to have relatively long legs compared with other harvestmen, and some species are gregarious.
Sabacon is a genus of the monotypic harvestman family Sabaconidae, with 59 species.
The Sironidae are a family of harvestmen with more than 60 described species.
Trogulidae is a family of harvestmen comprising 65 extant species in five genera.
Nipponopsalididae is a family of harvestmen with three described species in one genus, Nipponopsalis, which is found in East Asia.
Ischyropsalididae is a family of harvestmen with 35 described species in 3 genera, found in Europe and North America.
The Nemastomatidae are a family of harvestmen with about 170 described species in 16 recent genera. Several fossil species and genera are known.
The Dicranolasmatidae are a monotypic family of harvestmen with 16 described species in a single genus, Dicranolasma.
Cladolasma, is a genus of harvestmen belonging to the family Nemastomatidae. The genus is monotypic containing a single species Cladolasma parvulum. They are endemic to Japan, with records in Ehime and Tokushima Prefectures. The genus was described by Seisho Suzuki, with the type species Cladolasma parvulum Suzuki, 1963, subsequently revised to Dendrolasma parvulum in Suzuki (1974), but later restored as a valid genus.
Siro is a genus of mite harvestmen in the family Sironidae with 7 described species. All are found in Europe.
Nemastomatinae is a subfamily of harvestmen with over 125 described species in 19 (extant) genera. They are usually found in temperate regions.
Anelasmocephalus is a genus of harvestmen belonging to the family Trogulidae.
Asiolasma is a genus of harvestmen in the family Nemastomatidae with six described species . The species are found in Southeast Asia, namely China, Vietnam and Thailand. Some species are treated under other genera in Schönhofer (2013).
Troguloidea is a superfamily of harvestmen with 4 genera, found mostly in Europe, Asia, and North America.
Trogulus torosus is a species of large harvestman occurring in Boznia and Herzegovina, Croatia, and Montenegro. It is the largest harvestman in the world by body length, up to 23.2 mm.