Xenorhynchopsis Temporal range: | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Phoenicopteriformes |
Family: | Phoenicopteridae |
Genus: | † Xenorhynchopsis De Vis, 1905 |
Species | |
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Xenorhynchopsis is an extinct genus of flamingo from the Pliocene to Pleistocene Lake Eyre basin of Australia. Initially described as species of stork, the two known Xenorhynchopsis species are vastly different in size. X. minor is the older and small of the two species, ranging from the Pliocene to Pleistocene and being described as having reached a size smaller than that of the lesser flamingo. X. tibialis meanwhile appears to have been restricted to Pleistocene strata and was notably bigger, being counted as one of the biggest known flamingos in the fossil record.
The fossils of Xenorhynchopsis were initially described by Charles Walter de Vis in 1905, who recognized two species he named Xenorhynchopsis minor and Xenorhynchopsis tibialis respectively. Possibly due to the fact that de Vis lacked flamingo material in his collection to compare the fossils to, he assigned the genus to the Ciconiidae, the storks. X. tibialis, the larger of the two species, was described on the basis of a humerus and a tibiotarsus found in the Lower Cooper Creek. The smaller species X. minor meanwhile is known from multiple remains of the humerus and tibiotarsus discovered in Pliocene to Pleistocene sediments of Lake Kanunka and the Lower Cooper Creek. Rich and colleagues, who published a revision of the Pliocene and Pleistocene flamingo fossils of material, write that the remains are too fragmentary to determine whether or not these species should be placed in the same genera as modern flamingos. They subsequently retain the genus names coined by de Vis for the sake of convenience. [1] [2]
The traits that distinguish Xenorhynchopsis from other flamingo genera are deemed questionable by Rich and colleagues. The distal end of the tibiotarsus is described as deeper than it is wide with a shaft that flares less towards the distal end when compared to extant species. Towards the distal end the tendinal canal has larger openings, another trait shared by both species and setting them apart from modern flamingos. Additionally, the internal condyle lacks a notch in both species. [1]
The fossil material of X. tibialis resembles the black-necked stork in size and is larger than that of extant flamingos. Rich and colleagues write that only Phoeniconotius exceeds Xenorhynchopsis tibialis in size. X. minor meanwhile is somewhat smaller, if stouter, than the lesser flamingo and approximately in the same size range as Phoenicopterus stocki and Phoenicopterus minutus . [1]
During the Pliocene at least three flamingo species existed at Lake Kanunka, Xenorhynchopsis minor, Phoeniconaias proeses and what may be the modern greater flamingo. Xenorhynchopsis tibialis appears to have been limited to the Pleistocene and possibly died out when the inland lakes these birds depended on dried up, leading to the local extinction of flamingos in Australia. All four species could be found in the sediments of Lower Cooper Creek, however due to them appearing in different localities, many of which lack precise information regarding their age, it is uncertain if they actually occurred alongside one another. [1] According to Camens and Worthy, footprints found in the Tirari Formation may have been left by Xenorhynchopsis minor, as they appear to be too large to be attributed to P. proeses and too small to have belonged to X. tibialis. The greater flamingo is however another possible candidate. [3]
Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas, and two species native to Afro-Eurasia.
The darters, anhingas, or snakebirds are mainly tropical waterbirds in the family Anhingidae, which contains a single genus, Anhinga. There are four living species, three of which are very common and widespread while the fourth is rarer and classified as near-threatened by the IUCN. The term snakebird is usually used without any additions to signify whichever of the completely allopatric species occurs in any one region. It refers to their long thin neck, which has a snake-like appearance when they swim with their bodies submerged, or when mated pairs twist it during their bonding displays. "Darter" is used with a geographical term when referring to particular species. It alludes to their manner of procuring food, as they impale fishes with their thin, pointed beak. The American darter is more commonly known as the anhinga. It is sometimes called "water turkey" in the southern United States; though the anhinga is quite unrelated to the wild turkey, they are both large, blackish birds with long tails that are sometimes hunted for food.
The lesser flamingo is a species of flamingo occurring in sub-Saharan Africa and western India. Birds are occasionally reported from further north, but these are generally considered vagrants.
Moorhens—sometimes called marsh hens—are medium-sized water birds that are members of the rail family (Rallidae). Most species are placed in the genus Gallinula, Latin for "little hen". They are close relatives of coots. They are often referred to as (black) gallinules. Recently, one of the species of Gallinula was found to have enough differences to form a new genus Paragallinula with the only species being the lesser moorhen.
Ciconia is a genus of birds in the stork family. Six of the seven living species occur in the Old World, but the maguari stork has a South American range. In addition, fossils suggest that Ciconia storks were somewhat more common in the tropical Americas in prehistoric times.
Palaelodus is an extinct genus of bird of the Palaelodidae family, distantly related to flamingos. They were slender birds with long, thin legs and a long neck resembling their modern relatives, but likely lived very different livestyles. They had straight, conical beaks not suited for filter feeding and legs showing some similarities to grebes. Their precise lifestyle is disputed, with researchers in the past suggesting they may have been divers, while more recent research suggests they may have used their stiff toes as paddles for swimming while feeding on insect larvae and snails. This behavior may have been key in later phoenicopteriforms developing filterfeeding bills. The genus includes between five to eight species and is found across Europe, Australia, New Zealand, Asia and possibly South America. However some argue that most of the taxa named from Europe simply represent differently sized individuals of one single species. Palaelodus was most abundant during the Late Oligocene to Middle Miocene periods, but isolated remains from Australia indicate that the genus, or at least a relative, survived until the Pleistocene.
Megapaloelodus is an extinct genus of stem flamingo of the family Palaelodidae. Megapaloelodus is primarily known from Miocene America, from South Dakota and Oregon in the north to Argentina in the south, but the species Megapaloelodus goliath was found in Europe. Additionally, one unnamed species was discovered in Miocene sediments from Namibia. Due to a lack of skull material, little can be said about the ecology of Megapaloelodus. Species of this genus are typically larger than those of Palaelodus and appear to have inhabited similar brackish lake environments. Additionally, they may have been capable of "locking" their legs in a standing position.
Palaelodidae is a family of extinct birds in the group Phoenicopteriformes, which today is represented only by the flamingos. They were widespread during the Neogene, with fossil remains found on all continents other than Antarctica. The oldest remains referred to this group appeared in the fossil record during the Oligocene in Egypt and Belgium, before palaelodids reached their peak diversity during the Miocene. Following this the group declined in the early Pliocene before going extinct on most continents. However, remains found near Cooper Creek in the Lake Eyre Basin indicate that palaelodids managed to survive in Australia until the Pleistocene. Currently three genera are recognized by scientists: Adelalopus, Palaelodus and Megapaloelodus. Most fossil remains stem from Europe and have been assigned to the type species, Palaelodus ambiguus. Due to the fragmentary nature of most of these species, little is known about their ecology. They appear to have preferred brackish lakes and lagoons. Palaelodus has previously been thought to be a wader or diver, but recent research indicates that they were better suited for swimming and possibly fed on insect larvae and other aquatic invertebrates. At least Megapaloelodus appears to have adaptations for "locking" their legs in a standing position.
Leptoptilos robustus is an extinct species of large-bodied stork belonging to the genus Leptoptilos that lived on the island of Flores in Indonesia during the Pleistocene epoch. It stood at about 1.8 metres (5.9 ft) tall and weighed up to an estimated 16 kilograms (35 lb). The majority of the discoveries are concentrated in Liang Bua cave located slightly north of Ruteng in the East Nusa Tenggara province.
Phoeniconaias is a genus of birds in the flamingo family Phoenicopteridae. The genus contains one extant species, the lesser flamingo occurring in sub-Saharan Africa and western India, and an extinct species Phoeniconaias proeses from the Pliocene of Australia which is thought to have been even smaller.
Phoenicopterus minutus is an extinct species of flamingo which inhabited California during the Late Pleistocene. It was originally discovered in San Bernardino County, California in the Lake Manix beds, where it coexisted with a second, larger flamingo species.
Phoenicopterus copei is an extinct species of flamingo that inhabited North America during the Late Pleistocene. Its fossils have been discovered in Oregon, California, Mexico and Florida. Many of these localities preserve the remains of juvenile individuals, indicating that this species nested at the lakes found there. In some areas like California and Florida it coexisted with smaller flamingo species. P. copei was a large species of Phoenicopterus, described as being greater in size than modern American flamingos.
Cryptogyps is an extinct genus of Old World vulture from the Pleistocene of Australia. It was relatively small for a vulture but still larger than the extant wedge-tailed eagle. Originally described as an eagle in 1905, in 2022 it was reidentified as a vulture, the only known representative from the continent. Phylogenetic analysis supports it being a sister species to the extant, widespread Eurasian vulture genus Gyps or as a more basal member of the subfamily. The identification of Cryptogyps as a vulture solves a longstanding mystery about the lack of specialized lineages of large scavenging birds in Australia despite being present on every other continent aside from Antarctica. It is likely that Cryptogyps went extinct towards the end of the Pleistocene due to the disappearance of the megafauna it depended on for carrion. The genus contains a single species, C. lacertosus.
Leakeyornis is an extinct genus of flamingo from the early to middle Miocene of Kenya, primarily in the area of modern day Lake Victoria. Initially described as a species of Phoenicopterus based on an incomplete skull and various limb bones, it was later found to show a mixture of traits found across modern flamingo genera and subsequently placed in its own genus. It contains a single species, Leakeyornis aethiopicus.
Phoenicopterus novaehollandiae is an extinct species of flamingo from the late Oligocene or early Miocene Etadunna Formation of Australia. It was a large species similar in size to large specimens of the modern greater flamingo, but differs by likely having had a much better developed hallux which is typically reduced or absent in modern flamingos.
Phoenicopterus stocki, also known as Stock's flamingo, is an extinct species of flamingo from the Pliocene of Chihuahua, Mexico. It was described in 1944 as a small bodied flamingo species known from assorted fragmentary remains, including bones of the tibia and the wings. The discovery of juvenile remains suggests that the locality where the fossils were found represents a shallow lagoon or mudflat that housed a breeding colony.
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Pliolymbus is a fossil genus of grebe known from the Late Pliocene to Early Pleistocene of United States and Mexico. It is known from a single species, P. baryosteus.
Aechmophorus elasson is an extinct species of grebe recovered from the Piacenzian age of the United States.