Phoenicopterus stocki Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Phoenicopteriformes |
Family: | Phoenicopteridae |
Genus: | Phoenicopterus |
Species: | †P. stocki |
Binomial name | |
†Phoenicopterus stocki Miller, 1944 | |
Phoenicopterus stocki, also known as Stock's flamingo, is an extinct species of flamingo from the Pliocene of Chihuahua, Mexico. It was described in 1944 as a small bodied flamingo species known from assorted fragmentary remains, including bones of the tibia and the wings. The discovery of juvenile remains suggests that the locality where the fossils were found represents a shallow lagoon or mudflat that housed a breeding colony.
The remains of Phoenicopterus stocki were discovered in the early 20th century by a field party of the California Institute of Technology led by Chester Stock in the Mexican state of Chihuahua. The field party uncovered nine fragments belong to a species of flamingo later described by Loye H. Miller. The type specimen of this taxon was the distal end of a left tibiotarsus, but additional material from various parts of the body have also been referred to this species. This additional material includes the proximal end of a tibia, possibly belonging to the same individual as the holotype as well as two differently sized humerus fragments. Some of the material is thought to have belonged to juvenile birds not yet capable of flight. [1] All the material initially assigned to the species was uncovered from an area known as Arroyo de los Ponos, also known as CIT locality 289. Additional material would later be collected from the same site as well as the nearby Arroyo de los Burros. [2]
Miller described that Phoenicopterus stocki shares the morphology of extant Phoenicopterus species, but notes that the animal was much smaller, referring to it as a pygmy. The tibiotarsus generally agrees more with the American flamingo in its proportions, being shallower than in the greater flamingo. The intercondylar area is much more narrow than in any extant flamingo and the intercondylar notch, located on the elements distal border, is almost in the center of the condyle which sets it apart from the more recent Phoenicopterus minutus . [3] The fragment of the tibia is described as distinctive by Miller, but is not included in the species diagnosis due to the uncertain relationship between it and the holotype tibiotarsus. The inner articular facet for instance does not extend as far back as in either of the Neotropic Phoenicopterus species, the American and the Chilean flamingo, and instead appears to be closer to the greater flamingo of Africa and Eurasia. In the same fashion the postero-axial border of this facet is shaped almost like the arc of a circle and the posterior notch shallow, both characteristics shared by the greater flamingo but neither of the neotropic species. Although Miller did not have Phoenicoparrus skeletons available for comparison, he compared the Pliocene material to skins of said genus and determined that P. stocki was smaller than either James's flamingo or the Andean flamingo. Related to the tibia, Miller notes that the ratio between its transverse diameter and the sagittal diameter is greater than in the American flamingo but smaller than in Phoenicopterus copei . [1]
Both wing bones collected at the Rincón fossil locality are vastly different in size. While the larger is approximately the size of the same element in the Chilean flamingo, the smaller is outside the range of any adult flamingo known today. Miller places both specimens in P. stocki due to the great variability in leg to wing proportions, specifically citing Chilean flamingos and greater flamingos. Still, the possibility of a second flamingo from the locality is not discarded entirely. [1]
A later publication by Hildegarde Howard also mentioned the small size of Stock's flamingo, writing that the species was small in size, but not as small as Phoenicopterus minutus from the Pleistocene of California. [4]
Phoenicopterus stocki likely led a similar lifestyle to modern flamingos, wading through water and filtering small organisms using a specially adapted bill. Subsequently, it is inferred that the Rincón locality where the material has been found represents what used to be a shallow lagoon or mudflat during the Pliocene, a conclusion further supported by the presence of not yet volant juveniles. This would mean that flamingos not only fed at the side, but nested and raised their young there as well. [1] Support for this hypothesis is also lend by the other bird remains found there, which generally belong to aquatic species. [2]
Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas, and two species native to Afro-Eurasia.
The greater flamingo is the most widespread and largest species of the flamingo family. Common in the Old World, they are found in Northern (coastal) and Sub-Saharan Africa, the Indian Subcontinent, the Middle East, the Levant, the Persian Gulf, the Gulf of Aden, the Red Sea, and the Mediterranean countries of Southern Europe.
Ciconia is a genus of birds in the stork family. Six of the seven living species occur in the Old World, but the maguari stork has a South American range. In addition, fossils suggest that Ciconia storks were somewhat more common in the tropical Americas in prehistoric times.
Palaelodus is an extinct genus of bird of the Palaelodidae family, distantly related to flamingos. They were slender birds with long, thin legs and a long neck resembling their modern relatives, but likely lived very different livestyles. They had straight, conical beaks not suited for filter feeding and legs showing some similarities to grebes. Their precise lifestyle is disputed, with researchers in the past suggesting they may have been divers, while more recent research suggests they may have used their stiff toes as paddles for swimming while feeding on insect larvae and snails. This behavior may have been key in later phoenicopteriforms developing filterfeeding bills. The genus includes between five and eight species and is found across Europe, Australia, New Zealand, Asia and possibly South America. However some argue that most of the taxa named from Europe simply represent differently sized individuals of one single species. Palaelodus was most abundant during the Late Oligocene to Middle Miocene periods, but isolated remains from Australia indicate that the genus, or at least a relative, survived until the Pleistocene.
Megapaloelodus is an extinct genus of stem flamingo of the family Palaelodidae. Megapaloelodus is primarily known from Miocene America, from South Dakota and Oregon in the north to Argentina in the south, but the species Megapaloelodus goliath was found in Europe. Additionally, one unnamed species was discovered in Miocene sediments from Namibia. Due to a lack of skull material, little can be said about the ecology of Megapaloelodus. Species of this genus are typically larger than those of Palaelodus and appear to have inhabited similar brackish lake environments. Additionally, they may have been capable of "locking" their legs in a standing position.
Palaelodidae is a family of extinct birds in the group Phoenicopteriformes, which today is represented only by the flamingos. They were widespread during the Neogene, with fossil remains found on all continents other than Antarctica. The oldest remains referred to this group appeared in the fossil record during the Oligocene in Egypt and Belgium, before palaelodids reached their peak diversity during the Miocene. Following this the group declined in the early Pliocene before going extinct on most continents. However, remains found near Cooper Creek in the Lake Eyre Basin indicate that palaelodids managed to survive in Australia until the Pleistocene. Currently three genera are recognized by scientists: Adelalopus, Palaelodus and Megapaloelodus. Most fossil remains stem from Europe and have been assigned to the type species, Palaelodus ambiguus. Due to the fragmentary nature of most of these species, little is known about their ecology. They appear to have preferred brackish lakes and lagoons. Palaelodus has previously been thought to be a wader or diver, but recent research indicates that they were better suited for swimming and possibly fed on insect larvae and other aquatic invertebrates. At least Megapaloelodus appears to have adaptations for "locking" their legs in a standing position.
Leptoptilos robustus is an extinct species of large-bodied stork belonging to the genus Leptoptilos that lived on the island of Flores in Indonesia during the Pleistocene epoch. It stood at about 1.8 metres (5.9 ft) tall and weighed up to an estimated 16 kilograms (35 lb). The majority of the discoveries are concentrated in Liang Bua cave located slightly north of Ruteng in the East Nusa Tenggara province.
Phoenicopterus minutus is an extinct species of flamingo which inhabited California during the Late Pleistocene. It was originally discovered in San Bernardino County, California in the Lake Manix beds, where it coexisted with a second, larger flamingo species.
Phoenicopterus copei is an extinct species of flamingo that inhabited North America during the Late Pleistocene. Its fossils have been discovered in Oregon, California, Mexico and Florida. Many of these localities preserve the remains of juvenile individuals, indicating that this species nested at the lakes found there. In some areas like California and Florida it coexisted with smaller flamingo species. P. copei was a large species of Phoenicopterus, described as being greater in size than modern American flamingos.
Garganornis is an extinct genus of enormous flightless anatid waterfowl from the Late Miocene of Gargano, Italy. The genus contains one species, G. ballmanni, named by Meijer in 2014. Its enormous size is thought to have been an adaptation to living in exposed, open areas with no terrestrial predators, and as a deterrent to the indigenous aerial predators like the eagle Garganoaetus and the giant barn owl Tyto gigantea.
Chester Stock was an American paleontologist who specialized in the Pleistocene mammalian fauna of the Rancho La Brea tar pits. He served as a professor of geology at the California Institute of Technology, Pasadena.
Harrisonavis is an extinct genus of flamingo that lived during the Late Oligocene and Early Miocene periods in what is now France. Despite being one of the oldest known members of the flamingo family, it already shows a skull remarkably similar to that of the modern greater flamingo. Although generally similar, it subtly differs in the curvature of the bill and the size of the ventral keel of the maxilla, both signs that Harrisonavis was not yet as adapted towards filter feeding as modern species are. Harrisonavis inhabited brackish lakes alongside the more basal Palaelodidae. It contains the single species Harrisonavis croizeti, first described in 1852.
Asio ecuadoriensis is an extinct species of eared owl from the Pleistocene of Ecuador. Known from bones of the legs, it was a robust predatory bird similar in size to the great horned owl. Based on the robustness of its limbs and the bones found in what may have been its burrow, it's possible that it may have been specialised in hunting other species of owls.
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Leakeyornis is an extinct genus of flamingo from the early to middle Miocene of Kenya, primarily in the area of modern day Lake Victoria. Initially described as a species of Phoenicopterus based on an incomplete skull and various limb bones, it was later found to show a mixture of traits found across modern flamingo genera and subsequently placed in its own genus. It contains a single species, Leakeyornis aethiopicus.
Phoenicopterus novaehollandiae is an extinct species of flamingo from the late Oligocene or early Miocene Etadunna Formation of Australia. It was a large species similar in size to large specimens of the modern greater flamingo, but differed by likely having had a much better developed hallux which is typically reduced or absent in modern flamingos.
Phoenicopterus floridanus is an extinct species of flamingo that lived during the Pliocene in what is now Florida and potentially North Carolina.
Phoeniconaias proeses is an extinct species of flamingo from the Pliocene of Australia. Fossil material was described under several names including Ocyplanus proeses and Phoeniconaias gracilis, which were eventually found to be synonymous. Only material from the Tirari Formation has been dated, while most other material lacks precise information on its age. P. proeses was one of the smallest species of flamingo, smaller than the modern lesser flamingo which it may be related to.
Xenorhynchopsis is an extinct genus of flamingo from the Pliocene to Pleistocene Lake Eyre basin of Australia. Initially described as species of stork, the two known Xenorhynchopsis species are vastly different in size. X. minor is the older and small of the two species, ranging from the Pliocene to Pleistocene and being described as having reached a size smaller than that of the lesser flamingo. X. tibialis meanwhile appears to have been restricted to Pleistocene strata and was notably bigger, being counted as one of the biggest known flamingos in the fossil record.
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