Phoenicopterus minutus Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Phoenicopteriformes |
Family: | Phoenicopteridae |
Genus: | Phoenicopterus |
Species: | †P. minutus |
Binomial name | |
†Phoenicopterus minutus Howard, 1955 | |
Phoenicopterus minutus is an extinct species of flamingo which inhabited California during the Late Pleistocene. It was originally discovered in San Bernardino County, California in the Lake Manix beds, where it coexisted with a second, larger flamingo species. [1] [2] [3]
The fossils of Phoenicopterus minutus were discovered in the Lake Manix beds during a large scale project conducted by the United States Geological Survey. Although this project had begun in 1947, much of the bird material from Lake Manix was not collected until the 1950s. The material that formed the basis of the taxon consists of a nearly complete tibiotarsus alongside associated tarsometatarsus elements, which accounts for approximately two thirds of said bone. An additional tarsometatarsus fragment is also known, as are two scapulae and a fragment of the coracoid. The species was named by Hildegard Howard in 1955. [1]
Phoenicopterus minutus was said to be the smallest species of flamingo at the time of its description. Specifically, it was stated that the tibiotarsus was 13 mm (0.51 in) shorter than the same element in the smallest available Chilean flamingo skeleton. The distal end was also narrower and more shallow than in Phoenicopterus stocki from middle Pliocene Mexico. These two fossil forms further differ in their anatomy. Whereas the older P. stocki resembles the proportions of the modern American flamingo, P. minutus's shallower tibiotarsus bears closer resemblance to the greater flamingo. The notch present on the internal condyle is central in P. stocki, but moved well behind this point in P. minutus, a condition also seen in the American flamingo. While the tarsometatarsus could not be compared between these two species, there were still differences to extant forms such as the intercotylar tubercles, which are high and narrow. The ratio between the tubercles height to the width is noted to be even bigger than in American flamingos. Both scapula are similar in size but differ in their contour. This however is not overly exceptional as this also occurs in the still living flamingo species. More notable is the shape of the glenoid facet, the part of the shoulder that connects to the upper arms, which is inflated but smaller than in modern species. The coracoid, although larger than would be expected based on the type material, was deemed too incomplete form the basis of any further conclusions. [1]
Although it was initially described as one of the smallest known flamingos, later research uncovered the presence of even smaller species. According to Rich et al., Phoenicopterus minutus was in a similar size range as Xenorhynchopsis minor and Phoenicopterus stocki . They consider Phoeniconaias proeses and Leakeyornis to have been smaller. [4]
Lake Manix is known for the preservation of a multitude of water birds that lived in the region during the Pleistocene, including a second species of flamingo. This second species, known from juvenile remains that fall within the size range of modern American and greater flamingos has been tentatively assigned to Phoenicopterus copei . In general, the deposits of Lake Manix resemble that of Fossil Lake (Oregon), with a diverse avifauna including pelicans, grebes, cormorants, geese and ducks. [1]
Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas, and two species native to Afro-Eurasia.
Ciconia is a genus of birds in the stork family. Six of the seven living species occur in the Old World, but the maguari stork has a South American range. In addition, fossils suggest that Ciconia storks were somewhat more common in the tropical Americas in prehistoric times.
Palaelodus is an extinct genus of bird of the Palaelodidae family, distantly related to flamingos. They were slender birds with long, thin legs and a long neck resembling their modern relatives, but likely lived very different livestyles. They had straight, conical beaks not suited for filter feeding and legs showing some similarities to grebes. Their precise lifestyle is disputed, with researchers in the past suggesting they may have been divers, while more recent research suggests they may have used their stiff toes as paddles for swimming while feeding on insect larvae and snails. This behavior may have been key in later phoenicopteriforms developing filterfeeding bills. The genus includes between five and eight species and is found across Europe, Australia, New Zealand, Asia and possibly South America. However some argue that most of the taxa named from Europe simply represent differently sized individuals of one single species. Palaelodus was most abundant during the Late Oligocene to Middle Miocene periods, but isolated remains from Australia indicate that the genus, or at least a relative, survived until the Pleistocene.
Megapaloelodus is an extinct genus of stem flamingo of the family Palaelodidae. Megapaloelodus is primarily known from Miocene America, from South Dakota and Oregon in the north to Argentina in the south, but the species Megapaloelodus goliath was found in Europe. Additionally, one unnamed species was discovered in Miocene sediments from Namibia. Due to a lack of skull material, little can be said about the ecology of Megapaloelodus. Species of this genus are typically larger than those of Palaelodus and appear to have inhabited similar brackish lake environments. Additionally, they may have been capable of "locking" their legs in a standing position.
Phoeniconotius is an extinct genus of flamingo that lived in Australia from the late Oligocene to the early Miocene. Unlike modern flamingos and the contemporary Phoenicopterus novaehollandiae, it was likely less well adapted for swimming and deep water wading. Phoeniconotius was a robust flamingo with bones more massive than those of the modern greater flamingo. Only a single species is recognized, Phoeniconotius eyrensis.
Lake Manix is a former lake fed by the Mojave River in the Mojave Desert. It lies within San Bernardino County, California. Located close to Barstow, this lake had the shape of a cloverleaf and covered four basins named Coyote, Cady/Manix, Troy and Afton. It covered a surface area of 236 square kilometres (91 sq mi) and reached an altitude of 543 metres (1,781 ft) at highstands, although poorly recognizable shorelines have been found at altitudes of 547–558 metres (1,795–1,831 ft). The lake was fed by increased runoff during the Pleistocene and overflowed into the Lake Mojave basin and from there to Lake Manly in Death Valley, or less likely into the Bristol Lake basin and from there to the Colorado River.
Palaelodidae is a family of extinct birds in the group Phoenicopteriformes, which today is represented only by the flamingos. They were widespread during the Neogene, with fossil remains found on all continents other than Antarctica. The oldest remains referred to this group appeared in the fossil record during the Oligocene in Egypt and Belgium, before palaelodids reached their peak diversity during the Miocene. Following this the group declined in the early Pliocene before going extinct on most continents. However, remains found near Cooper Creek in the Lake Eyre Basin indicate that palaelodids managed to survive in Australia until the Pleistocene. Currently three genera are recognized by scientists: Adelalopus, Palaelodus and Megapaloelodus. Most fossil remains stem from Europe and have been assigned to the type species, Palaelodus ambiguus. Due to the fragmentary nature of most of these species, little is known about their ecology. They appear to have preferred brackish lakes and lagoons. Palaelodus has previously been thought to be a wader or diver, but recent research indicates that they were better suited for swimming and possibly fed on insect larvae and other aquatic invertebrates. At least Megapaloelodus appears to have adaptations for "locking" their legs in a standing position.
Phoenicopterus copei is an extinct species of flamingo that inhabited North America during the Late Pleistocene. Its fossils have been discovered in Oregon, California, Mexico and Florida. Many of these localities preserve the remains of juvenile individuals, indicating that this species nested at the lakes found there. In some areas like California and Florida it coexisted with smaller flamingo species. P. copei was a large species of Phoenicopterus, described as being greater in size than modern American flamingos.
Spheniscus muizoni is an extinct species of banded penguins that lived during the early Late Miocene in what is now Peru, South America. The species, the earliest member of the extant genus, was described in 2007 by Ursula B. Göhlich based on fossils found in the fossiliferous Pisco Formation of the Pisco Basin, southwestern Peru.
Harrisonavis is an extinct genus of flamingo that lived during the Late Oligocene and Early Miocene periods in what is now France. Despite being one of the oldest known members of the flamingo family, it already shows a skull remarkably similar to that of the modern greater flamingo. Although generally similar, it subtly differs in the curvature of the bill and the size of the ventral keel of the maxilla, both signs that Harrisonavis was not yet as adapted towards filter feeding as modern species are. Harrisonavis inhabited brackish lakes alongside the more basal Palaelodidae. It contains the single species Harrisonavis croizeti, first described in 1852.
Melanerpes shawi is an extinct species of woodpecker from the Pleistocene of California. It was found in the La Brea tar pits. It's part of the genus Melanerpes, which includes twenty-four extant species found across North and South America.
Leakeyornis is an extinct genus of flamingo from the early to middle Miocene of Kenya, primarily in the area of modern day Lake Victoria. Initially described as a species of Phoenicopterus based on an incomplete skull and various limb bones, it was later found to show a mixture of traits found across modern flamingo genera and subsequently placed in its own genus. It contains a single species, Leakeyornis aethiopicus.
Phoenicopterus novaehollandiae is an extinct species of flamingo from the late Oligocene or early Miocene Etadunna Formation of Australia. It was a large species similar in size to large specimens of the modern greater flamingo, but differed by likely having had a much better developed hallux which is typically reduced or absent in modern flamingos.
Phoenicopterus stocki, also known as Stock's flamingo, is an extinct species of flamingo from the Pliocene of Chihuahua, Mexico. It was described in 1944 as a small bodied flamingo species known from assorted fragmentary remains, including bones of the tibia and the wings. The discovery of juvenile remains suggests that the locality where the fossils were found represents a shallow lagoon or mudflat that housed a breeding colony.
Phoenicopterus floridanus is an extinct species of flamingo that lived during the Pliocene in what is now Florida and potentially North Carolina.
Phoeniconaias proeses is an extinct species of flamingo from the Pliocene of Australia. Fossil material was described under several names including Ocyplanus proeses and Phoeniconaias gracilis, which were eventually found to be synonymous. Only material from the Tirari Formation has been dated, while most other material lacks precise information on its age. P. proeses was one of the smallest species of flamingo, smaller than the modern lesser flamingo which it may be related to.
Xenorhynchopsis is an extinct genus of flamingo from the Pliocene to Pleistocene Lake Eyre basin of Australia. Initially described as species of stork, the two known Xenorhynchopsis species are vastly different in size. X. minor is the older and small of the two species, ranging from the Pliocene to Pleistocene and being described as having reached a size smaller than that of the lesser flamingo. X. tibialis meanwhile appears to have been restricted to Pleistocene strata and was notably bigger, being counted as one of the biggest known flamingos in the fossil record.
Pliolymbus is a fossil genus of grebe known from the Late Pliocene to Early Pleistocene of United States and Mexico. It is known from a single species, P. baryosteus.
Aechmophorus elasson is an extinct species of grebe recovered from the Piacenzian age of the United States.
Podiceps solidus is an extinct small species of Upper Miocene to Lower Pliocene grebe from Western Mongolia.