Aelurognathus

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Aelurognathus
Temporal range: Wuchiapingian, 260.9–254  Ma
Aelurognathus tigriceps.png
Holotype skull of A. tigriceps
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Gorgonopsia
Family: Gorgonopsidae
Subfamily: Rubidgeinae
Genus: Aelurognathus
Haughton, 1924
Type species
Scymnognathus tigriceps
Broom and Haughton, 1913
Synonyms

Genus-level

  • GorgonorhinusBroom, 1937
  • LeontocephalusBroom, 1940
  • ProrubidgeaBroom, 1940
  • TigricephalusBroom, 1948

Species-level

  • Scymnognathus serratidensHaughton, 1915
  • Gorgorhinus luckhoffiBroom, 1937
  • Leontocephalus cadleiBroom, 1940
  • Prorubidgea maccabeiBroom, 1940
  • Sycosaurus brodieiBroom, 1941
  • Gorgorhinus minorBroom, 1948
  • Tigricephalus kingwilliBroom, 1948
  • Lycaenops alticepsBrink & Kitching, 1953
  • Prorubidgea brinkiManten, 1959
  • Aelurognathus broodeiGebauer, 2007

Aelurognathus is an extinct genus of gorgonopsian therapsids from the Permian of South Africa and Zambia. [1]

Contents

Discovery

A. tigriceps restoration Aelur tigriceps1DB.jpg
A. tigriceps restoration
A. sp. skull at the Museum fur Naturkunde, Berlin Aelurognathus.jpg
A. sp. skull at the Museum für Naturkunde, Berlin

The type species is Aelurognathus tigriceps, originally named Scymnognathus tigriceps by South African paleontologists Robert Broom and Sydney H. Haughton in 1913, and later assigned to the new genus Aelurognathus by Haughton in 1924.

Scymnognathus parringtoni von Huene, 1950, previously assigned to Aelurognathus, is now classified as a species of Sauroctonus . [2] Aelurognathus nyasaensis Haughton, 1926 is not referable to the genus. [1]

Palaeobiology

A broken tooth beside the skeleton of a dicynodont from the Tropidostoma Assemblage Zone has been attributed to Aelurognathus, indicating that it scavenged. The bones of the back of the skeleton are the most scattered, suggesting that the Aelurognathus individuals fed on the rear of the carcass, removing the hind limbs to reach the soft underside. The small incisor teeth of Aelurognathus indicate that it was not able to crush bone but more likely stripped flesh from its prey like the modern-day wild dog Lycaon pictus . Bite marks on the bones of the skeleton were unlikely to have been made by Aelurognathus and may be an indication that another predator killed the dicynodont. [3]

Classification

Below is a cladogram from the phylogenetic analysis of Gebauer (2007): [4]

Restoration Leontocephalus.jpg
Restoration
Gorgonopsia 

Aloposaurus

Cyonosaurus

Aelurosaurus

Gorgonopsidae

Scylacognathus

Eoarctops

Gorgonops

Njalila

Lycaenops

Arctognathus

Inostrancevia

Rubidgeinae

Aelurognathus

Rubidgea

Sycosaurus

Clelandina

Related Research Articles

<span class="mw-page-title-main">Gorgonopsia</span> Extinct group of saber-toothed therapsids from the Permian

Gorgonopsia is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull, as well as elongated upper and sometimes lower canine teeth and incisors which were likely used as slashing and stabbing weapons. Postcanine teeth are generally reduced or absent. For hunting large prey, they possibly used a bite-and-retreat tactic, ambushing and taking a debilitating bite out of the target, and following it at a safe distance before its injuries exhausted it, whereupon the gorgonopsian would grapple the animal and deliver a killing bite. They would have had an exorbitant gape, possibly in excess of 90°, without having to unhinge the jaw.

<i>Gorgonops</i> Extinct genus of therapsids

Gorgonops is an extinct genus of gorgonopsian therapsid, of which it is the type genus. Gorgonops lived during the Late Permian (Wuchiapingian), about 260–254 million years ago in what is now South Africa.

<i>Inostrancevia</i> Extinct genus of therapsids

Inostrancevia is an extinct genus of large carnivorous therapsids which lived during the Late Permian in what is now European Russia and South Africa. The first known fossils of this gorgonopsian were discovered in the Northern Dvina, where two almost complete skeletons were exhumed. Subsequently, several other fossil materials were discovered in various oblasts, and these finds will lead to a confusion about the exact number of valid species in the country, before only three of them were officially recognized : I. alexandri, I. latifrons and I. uralensis. More recent research carried out in South Africa has discovered fairly well-preserved remains of the genus, being attributed to the species I. africana. The whole genus is named in honor of Alexander Inostrantsev, professor of Vladimir P. Amalitsky, the paleontologist who described the taxon.

<i>Lycaenops</i> Extinct genus of therapsids

Lycaenops ("wolf-face") is a genus of carnivorous therapsids. It lived during the Middle Permian to the early Late Permian, about 260 mya, in what is now South Africa.

<i>Dinogorgon</i> Extinct genus of therapsids

Dinogorgon is a genus of gorgonopsid from the Late Permian of South Africa and Tanzania. The generic name Dinogorgon is derived from Greek, meaning "terrible gorgon", while its species name rubidgei is taken from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. The type species of the genus is D. rubidgei.

<i>Sauroctonus</i> Extinct genus of therapsids

Sauroctonus is an extinct genus of gorgonopsian therapsids who lived during the end of the Middle Permian in what is now European Russia. The first fossils, discovered in Tatarstan, were first considered to belong to a new species of the South African genus Arctognathus, named A. progressus in 1938. The taxon will be designated as such until 1940, when it will be assigned to the genus Inostrancevia by Ivan Yefremov, before being definitively classified in a separate genus erected by Alexey Bystrow in 1955. The most complete known fossils of S. progressus include cranial and postcranial elements currently all recorded in Tatarstan. These elements show that the animal is a rather medium-sized gorgonopsian.

<i>Arctops</i> Extinct genus of therapsids

Arctops is an extinct genus of gorgonopsian therapsids known from the Late Permian of South Africa. It measured up to 2 metres in length and its skull was 30 centimetres (12 in) long. The type species is Arctops willistoni. A second species, A. watsoni, may be synonymous with A. willistoni. A. kitchingi may be a third species of Arctops, but it was only tentatively assigned to the genus when it was first named. Both were formally synonymized with A. willistoni by Christian Kammerer in 2017.

<i>Arctognathus</i> Extinct genus of therapsids

Arctognathus is an extinct genus of gorgonopsids that throve during the Late Permian in the Karoo basin of what is now South Africa.

<i>Rubidgea</i> Extinct genus of therapsids

Rubidgea is a genus of gorgonopsid from the upper Permian of South Africa and Tanzania, containing the species Rubidgea atrox. The generic name Rubidgea is sometimes believed to be derived from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. However, this generic name was actually erected in honor of Rubidge's paternal grandfather, Sydney Rubidge, who was a renowned fossil hunter. Its species name atrox is derived from Latin, meaning “fierce, savage, terrible”. Rubidgea is part of the gorgonopsian subfamily Rubidgeinae, a derived group of large-bodied gorgonopsians restricted to the Late Permian (Lopingian). The subfamily Rubidgeinae first appeared in the Tropidostoma Assemblage Zone. They reached their highest diversity in the Cistecephalus and Daptocephalus assemblage zones of the Beaufort Group in South Africa.

<i>Cyonosaurus</i> Extinct genus of therapsids

Cyonosaurus is a genus of gorgonopsian therapsids from the late Permian and possibly early Triassic of South Africa. Cyonosaurus was 0.6 to 1.1 metres in length, with a skull 9 to 18 centimetres in length. The type species Cyonosaurus longiceps was named in 1937.

<i>Aloposaurus</i> Extinct genus of therapsids

Aloposaurus is an extinct genus of gorgonopsian therapsids from the Late Permian of South Africa. It was first named by Robert Broom in 1910, and contains the type species A. gracilis, and possibly a second species A. tenuis. This small gorgonopsid had a slender narrow skull only 12 centimetres (4.7 in) long, with a total body length of 60–70 cm (2.0–2.3 ft).

<i>Aelurosaurus</i> Extinct genus of therapsids

Aelurosaurus is a small, carnivorous, extinct genus of gorgonopsian therapsids from the Middle Permian to Late Permian of South Africa. It was discovered in the Karoo Basin of South Africa, and first named by Richard Owen in 1881. It was named so because it appeared to be an ancestor for cat-like marsupials, but not yet a mammal itself. It contains five species, A. felinus, A. whaitsi, A. polyodon, A. wilmanae, and A.? watermeyeri. A. felinus, the type species, is generally well described with established features, while the other four species are not due to their poorly preserved holotypes.

<i>Scylacops</i> Extinct genus of therapsids

Scylacops is an extinct genus of Gorgonopsia. It was first named by Broom in 1913, and contains two species, S. bigendens, and S. capensis. Its fossils have been found in South Africa and Zambia. It is believed to be closely related to the Gorgonopsian Sauroctonus progressus. Scylacops was a moderately sized Gorgonopsid.

Broomisaurus is an extinct genus of Gorgonopsia. It was first named by Joleaud in 1920, and contains the single species B. planiceps. Gebauer (2007) considered Broomisaurus to be a nomen dubium, indistinguishable as a separate taxon of gorgonopsian because it is based on only a fragmentary remains. A 2015 paper on Eriphostoma tentatively agreed with Gebauer's determination, but did not rule out the possibility that Broomisaurus might be synonymous with Eriphostoma.

Cerdorhinus is an extinct genus of gorgonopsian therapsids from the Permian of South Africa. The type species Cerdorhinus parvidens was named by South African paleontologist Robert Broom in 1936. A second species, Cerdorhinus rubidgei, was named in 1937. In 2007, a specimen of the latter was reassigned to the genus Cyonosaurus.

"Dixeya" nasuta is a species of the predatory gorgonopsian therapsids from the Late Permian of East Africa, known from fossils found in what is now Tanzania. The species has a complicated taxonomic history, it was originally named as a second species of the genus Dixeya which is now considered a junior synonym of Aelurognathus. "D." nasuta itself, however, was not moved to Aelurognathus, and although it was instead tentatively referred to Arctognathus at first it has since been recognised to not belong to this genus either. This situation leaves "Dixeya" nasuta without a formal genus name. It was proposed to belong to a new distinct genus, named "Njalila", that was informally proposed for the species in a PhD thesis, but this name has not yet been formally published and is currently a nomen nudum. "D." nasuta has been characterised from other gorgonopsians by a combination of its straight snout profile, upturned and 'pinched' nose, and curved jaw margin.

<i>Turfanodon</i> Extinct genus of dicynodonts

Turfanodon is an extinct genus of dicynodont therapsid from the Late Permian Sunan, Guodikeng, and Naobaogou Formations of China. The holotype of T. bogdaensis was discovered between 1963-1964 and was originally named in 1973 by A. Sun with the type species Turfanodon bogdaensis, Turfanodon was reclassified as a junior synonym of the related Dicynodon in 1988 by G. M. King. T. bogdaensis remained a species of Dicynodon for over two decades before the genus was reinstated in 2011 in a revision of the taxonomy of Dicynodon by palaeontologist Christian Kammerer. A second species from Inner Mongolia, T. jiufengensis, was named in 2021 by palaeontologist Jun Liu from a nearly complete skeleton and other referred bones. Turfanodon was a relatively large dicynodont, and similar in appearance to the related Daptocephalus from South Africa.

<i>Bulbasaurus</i> Extinct genus of dicynodonts

Bulbasaurus is an extinct genus of dicynodont that is known from the Lopingian epoch of the Late Permian period of what is now South Africa, containing the type and only species B. phylloxyron. It was formerly considered as belonging to Tropidostoma; however, due to numerous differences from Tropidostoma in terms of skull morphology and size, it has been reclassified the earliest known member of the family Geikiidae, and the only member of the group known from the Tropidostoma Assemblage Zone. Within the Geikiidae, it has been placed close to Aulacocephalodon, although a more basal position is not implausible.

Leogorgon is a extinct genus of dubious therapsid from the Late Permian Sokolki Faunal Assemblage of Russia. It was originally classified as a rubidgeine gorgonopsian, and would have been the first member of that clade from outside of Africa if that identification had been valid. However, it may instead be a combination of the tooth of a gorgonopsian and the braincase of a dicynodont, and may be a wastebin taxon.

Phorcys is an extinct genus of gorgonopsian that lived during the Middle Permian period (Guadalupian) of what is now South Africa. It is known from two specimens, both portions from the back of the skull, that were described and named in 2022 as a new genus and species P. dubei by Christian Kammerer and Bruce Rubidge. The generic name is from Phorcys of Greek mythology, the father of the Gorgons from which the gorgonopsians are named after, and refers to its status as one of the oldest representatives of the group in the fossil record. Phorcys was recovered from the lowest strata of the Tapinocephalus Assemblage Zone (AZ) of the Beaufort Group, making it one of the oldest known gorgonopsians in the fossil record—second only to fragmentary remains of an indeterminate gorgonopsian from the older underlying Eodicynodon Assemblage Zone.

References

  1. 1 2 Kammerer CF. (2016) Systematics of the Rubidgeinae (Therapsida: Gorgonopsia) PeerJ 4:e1608 https://doi.org/10.7717/peerj.1608
  2. Gebauer E I, 2014. Re-assessment of the taxonomic position of the specimen GPIT/RE/7113 (Sauroctonus parringtoni comb. nov., Gorgonopsia). In: Kammerer C F, Angielczyk K D, Fröbisch J eds. Early Evolutionary History of the Synapsida. Dordrecht: Springer. 185−207.
  3. Fordyce, N.; Smith, R.; Chinsamy, A. (2012). "Evidence of a therapsid scavenger in the Late Permian Karoo Basin, South Africa". South African Journal of Science. 108 (11/12). doi: 10.4102/sajs.v108i11/12.1158 . hdl: 11427/29098 .
  4. Gebauer, E.V.I. (2007). Phylogeny and evolution of the Gorgonopsia with a special reference to the skull and skeleton of GPIT/RE/7113 ("Aelurognathus"? parringtoni) (PDF) (Thesis). Dissertation Universität Tübingen.