Leogorgon

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Leogorgon
Temporal range: Late Permian, Vyatkian
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Leogorgon tooth.png
The tooth assigned to Leogorgon that probably belongs to a gorgonopsid
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Synapsida
Clade: Therapsida
Clade: Gorgonopsia (?)
Genus: Leogorgon
Ivakhnenko, 2003
Species:
L. klimovensis
Binomial name
Leogorgon klimovensis
Ivakhnenko, 2003

Leogorgon ("Leo's gorgon") is an extinct genus of dubious therapsid from the Late Permian Sokolki Faunal Assemblage of Russia. It was originally classified as a rubidgeine gorgonopsian, and would have been the first member of that clade from outside of Africa if that identification had been valid. However, it may instead be a combination of the tooth of a gorgonopsian and the braincase of a dicynodont, and may be a wastebin taxon.

History

Leogorgon klimovensis was named in 2003 by the paleontologist Mikhail Ivakhnenko. [1] The genus name honors the paleontologist Leonid Tatarinov. The holotype, PIN 4549/13, is a partial braincase from the Klimovo-1 locality in Vologda Oblast, Russia, and pertains to the Sokolki Faunal Assemblage. Ivakhnenko also referred an incisor tooth from the same locality to the new species. He interpreted the fossils as representing a large rubidgeine gorgonopsian, the first found outside of Africa, and noted that it was especially similar to Dinogorgon .

In 2008, however, Ivakhnenko noted that, due to its poorly known anatomy, Leogorgon could be a relative of the Russian Phthinosuchidae rather than the sole Russian representative of Rubidgeinae. [2] In 2016, a rubidgeine identity was formally rejected by paleontologist Christian Kammerer, as the features which Ivakhnenko stated supported its assignment to Rubidgeinae are actually present in other gorgonopsians, and in fact the braincase has no uniquely gorgonopsian features at all and may be from a dicynodont. [3] The referred incisor is from a gorgonopsian, but cannot be distinguished from incisors of the contemporaneous Inostrancevia . Recent evidence suggests that African gorgonopsians form an endemic group. [4]

Related Research Articles

<span class="mw-page-title-main">Gorgonopsia</span> Extinct group of saber-toothed therapsids from the Permian

Gorgonopsia is an extinct clade of sabre-toothed therapsids from the Middle to Upper Permian roughly 265 to 252 million years ago. They are characterised by a long and narrow skull, as well as elongated upper and sometimes lower canine teeth and incisors which were likely used as slashing and stabbing weapons. Postcanine teeth are generally reduced or absent. For hunting large prey, they possibly used a bite-and-retreat tactic, ambushing and taking a debilitating bite out of the target, and following it at a safe distance before its injuries exhausted it, whereupon the gorgonopsian would grapple the animal and deliver a killing bite. They would have had an exorbitant gape, possibly in excess of 90°, without having to unhinge the jaw.

<i>Inostrancevia</i> Extinct genus of therapsids

Inostrancevia is an extinct genus of large carnivorous therapsids which lived during the Late Permian in what is now European Russia and South Africa. The first known fossils of this gorgonopsian were discovered in the Northern Dvina, where two almost complete skeletons were exhumed. Subsequently, several other fossil materials were discovered in various oblasts, and these finds will lead to a confusion about the exact number of valid species in the country, before only three of them were officially recognized : I. alexandri, I. latifrons and I. uralensis. More recent research carried out in South Africa has discovered fairly well-preserved remains of the genus, being attributed to the species I. africana. The whole genus is named in honor of Alexander Inostrantsev, professor of Vladimir P. Amalitsky, the paleontologist who described the taxon.

<i>Dinogorgon</i> Extinct genus of therapsids

Dinogorgon is a genus of gorgonopsid from the Late Permian of South Africa and Tanzania. The generic name Dinogorgon is derived from Greek, meaning "terrible gorgon", while its species name rubidgei is taken from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. The type species of the genus is D. rubidgei.

<i>Daptocephalus</i> Assemblage Zone

The Daptocephalus Assemblage Zone is a tetrapod assemblage zone or biozone found in the Adelaide Subgroup of the Beaufort Group, a majorly fossiliferous and geologically important geological Group of the Karoo Supergroup in South Africa. This biozone has outcrops located in the upper Teekloof Formation west of 24°E, the majority of the Balfour Formation east of 24°E, and the Normandien Formation in the north. It has numerous localities which are spread out from Colesberg in the Northern Cape, Graaff-Reniet to Mthatha in the Eastern Cape, and from Bloemfontein to Harrismith in the Free State. The Daptocephalus Assemblage Zone is one of eight biozones found in the Beaufort Group and is considered Late Permian (Lopingian) in age. Its contact with the overlying Lystrosaurus Assemblage Zone marks the Permian-Triassic boundary.

<i>Sauroctonus</i> Extinct genus of therapsids

Sauroctonus is an extinct genus of gorgonopsian therapsids who lived during the end of the Middle Permian in what is now European Russia. The first fossils, discovered in Tatarstan, were first considered to belong to a new species of the South African genus Arctognathus, named A. progressus in 1938. The taxon will be designated as such until 1940, when it will be assigned to the genus Inostrancevia by Ivan Yefremov, before being definitively classified in a separate genus erected by Alexey Bystrow in 1955. The most complete known fossils of S. progressus include cranial and postcranial elements currently all recorded in Tatarstan. These elements show that the animal is a rather medium-sized gorgonopsian.

<i>Viatkogorgon</i> Extinct genus of therapsids

Viatkogorgon is a genus of gorgonopsian that lived during the Permian period in what is now Russia. The first fossil was found at the Kotelnich locality near the Vyatka River and was made the holotype of the new genus and species V. ivachnenkoi in 1999. The generic name refers to the river and the related genus Gorgonops—the gorgons of Greek mythology are often referenced in the names of the group. The specific name honors the paleontologist Mikhail F. Ivakhnenko. The holotype skeleton is one of the most complete gorgonopsian specimens known and includes rarely preserved elements such as gastralia and a sclerotic ring. A larger, but poorly preserved specimen has also been assigned to the species.

<i>Aelurognathus</i> Extinct genus of therapsids

Aelurognathus is an extinct genus of gorgonopsian therapsids from the Permian of South Africa and Zambia.

<i>Rubidgea</i> Extinct genus of therapsids

Rubidgea is a genus of gorgonopsid from the upper Permian of South Africa and Tanzania, containing the species Rubidgea atrox. The generic name Rubidgea is sometimes believed to be derived from the surname of renowned Karoo paleontologist, Professor Bruce Rubidge, who has contributed to much of the research conducted on therapsids of the Karoo Basin. However, this generic name was actually erected in honor of Rubidge's paternal grandfather, Sydney Rubidge, who was a renowned fossil hunter. Its species name atrox is derived from Latin, meaning “fierce, savage, terrible”. Rubidgea is part of the gorgonopsian subfamily Rubidgeinae, a derived group of large-bodied gorgonopsians restricted to the Late Permian (Lopingian). The subfamily Rubidgeinae first appeared in the Tropidostoma Assemblage Zone. They reached their highest diversity in the Cistecephalus and Daptocephalus assemblage zones of the Beaufort Group in South Africa.

<i>Clelandina</i> Extinct genus of therapsids

Clelandina is an extinct genus of rubidgeine gorgonopsian from the Late Permian of Cistecephalus Assemblage Zone of South Africa. It was first named by Broom in 1948. The type and only species is C. rubidgei. It is relatively rare, with only four known specimens.

<i>Pravoslavlevia</i> Extinct genus of therapsids

Pravoslavlevia is an extinct genus of gorgonopsian therapsids that lived in the late Permian and is part of the Sokolki subcomplex of Russia. It had a skull 22 centimetres (8.7 in) long. The total length of the animal was about 1.4 metres. Only one species is known.

<i>Megawhaitsia</i> Extinct genus of large therapsids from the Late Permian in East Russia

Megawhaitsia is an extinct genus of large therocephalian therapsids who lived during the Late Permian (Wuchiapingian) in what is now Eastern Europe. The only known species is M. patrichae, described in 2008 from several fossils discovered in various oblasts of European Russia. The fossils are representative of a large animal whose skull size is estimated to be 40–50 cm (16–20 in) long.

<i>Daptocephalus</i> Extinct genus of dicynodonts

Daptocephalus is an extinct genus of non-mammalian synapsid anomodont dicynodont, it which was found in Late Permian strata, in a biozone known precisely for the presence of fossils of this dicynodont, the Daptocephalus Assemblage Zone, in the Karoo Basin in South Africa. An additional species, D. huenei, is known from the Usili Formation in Tanzania and was formerly assigned to the genus Dicynodon before a study in 2019 recognised that the type specimen belonged to Daptocephalus.

<i>Eriphostoma</i> Extinct genus of therapsids

Eriphostoma is an extinct genus of gorgonopsian therapsids known from the Middle Permian of Tapinocephalus Assemblage Zone, South Africa. It has one known species, Eriphostoma microdon, and was first named by Robert Broom in 1911. It is the oldest known gorgonopsian and among the smallest and most basal members of the clade.

<i>Smilesaurus</i> Extinct genus of therapsids

Smilesaurus is an extinct genus of gorgonopsian known from Africa. It lived during the Late Permian. It contains the single species S. ferox.

Wangwusaurus is extinct genus of probable therapsid that lived in the Late Permian in present-day China. Only species is known, Wangwusaurus tayuensis, described by the paleontologist Yang Zhongjian in 1979 from seventeen teeth found in the Jiyuan formation, of which at least seven are recognized as not belonging to those of therapsids.

"Dixeya" nasuta is a species of the predatory gorgonopsian therapsids from the Late Permian of East Africa, known from fossils found in what is now Tanzania. The species has a complicated taxonomic history, it was originally named as a second species of the genus Dixeya which is now considered a junior synonym of Aelurognathus. "D." nasuta itself, however, was not moved to Aelurognathus, and although it was instead tentatively referred to Arctognathus at first it has since been recognised to not belong to this genus either. This situation leaves "Dixeya" nasuta without a formal genus name. It was proposed to belong to a new distinct genus, named "Njalila", that was informally proposed for the species in a PhD thesis, but this name has not yet been formally published and is currently a nomen nudum. "D." nasuta has been characterised from other gorgonopsians by a combination of its straight snout profile, upturned and 'pinched' nose, and curved jaw margin.

<span class="mw-page-title-main">Dicynodontoidea</span> Extinct infraorder of dicynodonts

Dicynodontoidea is an infraorder of dicynodont therapsids that includes the famous dicynodont Dicynodon, Lystrosaurus and the Triassic Kannemeyeriiformes, as well as numerous other closely related species. The name was coined by American paleontologist Everett C. Olson in 1941 as an infraorder, despite using the typical "-oidea" suffix of superfamilies, and was later redefined under a phylogenetic context in 2009 by paleontologist Christian F. Kammerer.

<span class="mw-page-title-main">Rubidgeinae</span> Extinct subfamily of mammals

Rubidgeinae is an extinct subfamily of gorgonopsid therapsids known only from Africa. They were among the largest gorgonopsians, and their fossils are common in the Cistecephalus and Daptocephalus assemblage zones of the Karoo Basin. They lived during the Late Permian, and became extinct at the end of the Permian.

<i>Nochnitsa</i> Extinct genus of therapsids

Nochnitsa is an extinct genus of gorgonopsian therapsids who lived during an uncertain stage of the Permian in what is now European Russia. Only one species is known, N. geminidens, described in 2018 from a single specimen including a complete skull and some postcranial remains, discovered in the red beds of Kotelnich, Kirov Oblast. The genus is named in reference to Nocnitsa, a nocturnal creature from Slavic mythology. This name is intended as a parallel to the Gorgons, which are named after many genera among gorgonopsians, as well as for the nocturnal behavior inferred for the animal. The only known specimen of Nochnitsa is one of the smallest gorgonopsians identified to date, with a skull measuring close to 8 cm (3.1 in) in length. The rare postcranial elements indicate that the animal's skeleton should be particularly slender.

Phorcys is an extinct genus of gorgonopsian that lived during the Middle Permian period (Guadalupian) of what is now South Africa. It is known from two specimens, both portions from the back of the skull, that were described and named in 2022 as a new genus and species P. dubei by Christian Kammerer and Bruce Rubidge. The generic name is from Phorcys of Greek mythology, the father of the Gorgons from which the gorgonopsians are named after, and refers to its status as one of the oldest representatives of the group in the fossil record. Phorcys was recovered from the lowest strata of the Tapinocephalus Assemblage Zone (AZ) of the Beaufort Group, making it one of the oldest known gorgonopsians in the fossil record—second only to fragmentary remains of an indeterminate gorgonopsian from the older underlying Eodicynodon Assemblage Zone.

References

  1. Ivakhnenko, M. F. (2003). "Eotherapsids from the East European Placket (Late Permian)". Paleontological Journal. 37: 339–465.
  2. Ivakhnenko, M. F. (2008). "Cranial morphology and evolution of Permian Dinomorpha (Eotherapsida) of Eastern Europe". Paleontological Journal. 42 (9): 859–995. doi:10.1134/S0031030108090013. S2CID   85114195.
  3. Kammerer, Christian F. (2016). "Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)". PeerJ. 4: e1608. doi: 10.7717/peerj.1608 . PMC   4730894 . PMID   26823998.
  4. Kammerer, Christian F.; Masyutin, Vladimir (2016). "Gorgonopsian therapsids (Nochnitsa gen. nov. and Viatkogorgon) from the Permian Kotelnich locality of Russia". PeerJ. 6: e4954. doi: 10.7717/peerj.4954 . PMC   5995105 . PMID   29900078.