Afromyrma Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Formicidae |
Subfamily: | Myrmicinae |
Tribe: | incertae sedis |
Genus: | † Afromyrma Dlussky, Brothers & Rasnitsyn, 2004 |
Species: | †A. petrosa |
Binomial name | |
†Afromyrma petrosa Dlussky, Brothers & Rasnitsyn, 2004 | |
Afromyrma is an extinct genus of ant in the formicid subfamily Myrmicinae, and is currently unplaced in any of the tribes of that subfamily. The genus contains a single described species, Afromyrma petrosa. Afromyrma is known from a single Upper Cretaceous fossil, which was found in Orapa in the Central District of Botswana. [1]
Afromyrma is known from a solitary fossil from Orapa, the holotype, specimen number BP/2/26501 tentatively identified as a female, though there is a possibility this is incorrect. [1] The holotype specimen is mostly complete but is rather poor in details, and is preserved as a compression fossil in shale. The shales and mudstones of the Orapa site were rapidly deposited in a crater lake formed as the result of a large volcanic eruption in the Cretaceous. The sediment originated as an epiclastic kimberlite and derives from rapid weathering and decomposition of the sides of the kimberlite crater. Of the four major sedimentation types that have been observe at Orapa, fossils are found in only two, the granular mass flows and the fine-grained sediments: woody debris fossils are found in the granular mass flows, but well-preserved fossils are only recovered in the fine-grained sediments. The fossil-bearing stratum was uncovered during mining of the underlying diamond-bearing rocks at the site. [2] Radiometric dating of the kimberlite pipe reported on in 1977 gave a general age of approximately 84 to 92 million years ago. [2] Further dating utilizing zircon crystals reported on in 2004 gave more defined age of approximately 91 million years ago placing the date in the Turonian age of the late Cretaceous. [1]
The fossil was first studied by Russian paleontologists Gennady M. Dlussky and Alexandr Rasnitsyn with South African paleontologist Denis Brothers. Their 2004 type description of the new genus and species was published in the journal Insect Systematics and Evolution . [1] The genus name Afromyrma is a combination of Africa, where the fossil was found, and the Greek myrmex which means "ant". The specific epithet petrosa, a derivation of the Latin word petra meaning rock is a reference to the fossil status of the species. [1]
The solitary Afromyrma specimen is incomplete and shows poor preservation in general with portions of the legs and antennae missing or very indistinct. [1] Overall it is estimated the full body would have been around 10.3 millimetres (0.41 in) long. Overall the shape of the head capsule is shorter than it is wide, with eyes located slightly to the rear of the heads midpoint and having an oval shape. The mandibles are triangular in outline with distinct teeth on the chewing margin. The specimen has a waist divided into two segments and the petiole shows a weakly developed node. The abdomen segmentation and general form of the abdominal apex are suggestive of a male ant, but the soft nature of the abdominal tissue makes this interpretation problematic, in contrast the harder head capsule is more reminiscent of a female and thus the fossil is interpreted as such. [1]
Gesomyrmex pulcher is an extinct species of ant in the subfamily Formicinae known from an Eocene fossil found in Europe. G. pulcher is one of only eight species in the ant genus Gesomyrmex to have been described from fossils found in Europe.
Ypresiomyrma is an extinct genus of ants in the subfamily Myrmeciinae that was described in 2006. There are four species described; one species is from the Isle of Fur in Denmark, two are from the McAbee Fossil Beds in British Columbia, Canada, and the fourth from the Bol’shaya Svetlovodnaya fossil site in Russia. The queens of this genus are large, the mandibles are elongated and the eyes are well developed; a stinger is also present. The behaviour of these ants would have been similar to that of extant Myrmeciinae ants, such as solitary foraging for arthropod prey and never leaving pheromone trails. The alates were poor flyers due to their size, and birds and animals most likely preyed on these ants. Ypresiomyrma is not assigned to any tribe, and is instead generally regarded as incertae sedis within Myrmeciinae. However, some authors believe Ypresiomyrma should be assigned as incertae sedis within Formicidae.
Afropone is an extinct genus of ant in the formicid subfamily Ponerinae, and is currently unplaced in any of the tribes of that subfamily. The genus contains two described species, Afropone oculata and Afropone (?) orapa. Afropone is known from a group of Upper Cretaceous fossils, which were found at Orapa in the Central District of Botswana.
Archimyrmex is an extinct genus of ant in the formicid subfamily Myrmeciinae, described by palaeoentomologist Theodore Cockerell in 1923. The genus contains four described species, Archimyrmex rostratus, Archimyrmex piatnitzkyi, Archimyrmex smekali and Archimyrmex wedmannae. Archimyrmex is known from a group of Middle Eocene fossils which were found in North America, South America, and Europe. The genus was initially placed in the subfamily Ponerinae, but it was later placed in Myrmeciinae; it is now believed to be the ancestor of the extant primitive genus Myrmecia from Australia. Despite this, Archimyrmex is not a member to any tribe and is regarded as incertae sedis within Myrmeciinae. However, some authors believe Archimyrmex should be assigned as incertae sedis within Formicidae. These ants can be characterised by their large mandibles and body length, ranging from 13.2 to 30 mm. They also have long, thin legs and an elongated mesosoma (thorax) and petiole.
Emplastus is an extinct morphogenus of ants in the subfamily Dolichoderinae, known from fossils found in Asia and Europe. The genus contains twelve species described from sites in England, Eastern Europe and Far Eastern Russia.
Casaleia is an extinct genus of ants in the formicid subfamily Amblyoponinae described by Pagliano & Scaramozzino in 1990 from fossils found in Europe. The genus contains four species dating from the Eocene to Miocene, Casaleia eocenica, Casaleia inversa, Casaleia longiventris, Casaleia orientalis.
Zherichinius is an extinct genus of ants in the subfamily Dolichoderinae known from fossils found in amber from the Middle Eocene of Sakhalin island Far eastern Russia and Bitterfeld, Germany. At the time of description the species Zherichinius horribilis and Zherichinius rapax were two of eight ant species known from Sakhalin fossils.
Pachycondyla eocenica is an extinct species of ant in the formicid subfamily Ponerinae described from fossils found in Europe. P. eocenica is one of six Lutetian Pachycondyla species.
Pachycondyla lutzi is an extinct species of ant in the formicid subfamily Ponerinae described by from fossils found in Europe. P. lutzi is one of six Lutetian Pachycondyla species.
Pachycondyla? messeliana is an extinct species of ants in the formicid subfamily Ponerinae described by from a fossil found in Europe. P.? messeliana is one of six Lutetian Pachycondyla species.
Pachycondyla parvula is an extinct species of ant in the formicid subfamily Ponerinae described by from a fossil found in Europe. P. parvula is one of six Lutetian Pachycondyla species.
Pachycondyla petiolosa is an extinct species of ant in the formicid subfamily Ponerinae described by from a fossil found in Europe. P. parvula is one of six Lutetian Pachycondyla species.
Pachycondyla petrosa is an extinct species of ant in the formicid subfamily Ponerinae described from a fossil found in Europe. P. petrosa is one of six Lutetian Pachycondyla species.
Liometopum incognitum is an extinct species of formicid in the ant subfamily Dolichoderinae known from fossils found in eastern Asia.
Aneuretellus is an extinct genus of ant in the formicid subfamily Aneuretinae, and is one of eight genera of the subfamily. The genus contains a single described species Aneuretellus deformis and is known from one Middle Eocene fossil which was found in Sakhalin in the Russian Far East.
Gesomyrmex macrops is an extinct species of formicid in the ant subfamily Formicinae known from a fossil found in eastern Asia.
Gesomyrmex incertus is an extinct species of formicid in the ant subfamily Formicinae known from a fossil found in eastern Asia.
Pachycondyla oligocenica is an extinct species of formicid in the ant subfamily Ponerinae known from a fossil found in eastern Asia.
Agastomyrma is an extinct genus of formicid in the ant subfamily Myrmicinae known from the fossil species Agastomyrma laticeps found in eastern Asia.
Electromyrmex is an extinct genus of ants in the formicid subfamily Dolichoderinae. The genus contains a single described species, Electromyrmex klebsi and is known from a group of Middle Eocene fossils which were found in Europe.