Beavers are large, semiaquatic rodents of the Northern Hemisphere. There are two existing species: the North American beaver and the Eurasian beaver. Beavers are the second-largest living rodents, after capybaras, weighing up to 50 kg (110 lb). They have stout bodies with large heads, long chisel-like incisors, brown or gray fur, hand-like front feet, webbed back feet, and tails that are flat and scaly. The two species differ in skull and tail shape and fur color. Beavers can be found in a number of freshwater habitats, such as rivers, streams, lakes and ponds. They are herbivorous, consuming tree bark, aquatic plants, grasses and sedges.
The Oligocene is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present. As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from Ancient Greek ὀλίγος (olígos) 'few' and καινός (kainós) 'new', and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period.
Rails are a large, cosmopolitan family of small- to medium-sized terrestrial and/or semi-amphibious birds. The family exhibits considerable diversity in its forms, and includes such ubiquitous species as the crakes, coots, and gallinule; other rail species are extremely rare or endangered. Many are associated with wetland habitats, some being semi-aquatic like waterfowl, but many more are wading birds or shorebirds. The ideal rail habitats are marsh areas, including rice paddies, and flooded fields or open forest. They are especially fond of dense vegetation for nesting. The rail family is found in every terrestrial habitat with the exception of dry desert, polar or freezing regions, and alpine areas. Members of Rallidae occur on every continent except Antarctica. Numerous unique island species are known.
Castoridae is a family of rodents that contains the two living species of beavers and their fossil relatives. A formerly diverse group, only a single genus is extant today, Castor. Two other genera of "giant beavers", Castoroides and Trogontherium, became extinct in the Late Pleistocene.
Sciuromorpha ( 'squirrel-like') is a rodent clade that includes several different rodent families. It includes all members of the Sciuridae as well as the mountain beaver species.
Castorimorpha is the suborder of rodents containing the beavers and the kangaroo rats. A 2017 study using retroposon markers indicated that they are most closely related to the Anomaluromorpha and Myomorpha.
Palaeocastor is an extinct genus of beavers that lived in the North American Badlands during the late Oligocene period to early Miocene. Palaeocastor was much smaller than modern beavers. There are several species including Palaeocastor fossor, Palaeocastor magnus,Palaeocastor wahlerti, and Palaeocastor peninsulatus. The animals first became known on grounds of their fossilized burrows, the "Devil's corkscrews".
Daeodon is an extinct genus of entelodont even-toed ungulates that inhabited North America about 23 to 20 million years ago during the latest Oligocene and earliest Miocene. The type species is Daeodon shoshonensis, described by a very questionable holotype by Cope. Some authors synonymize it with Dinohyus hollandi and several other species, but due to the lack of diagnostic material, this is questionable at best.
Steneofiber is an extinct genus of beavers from the Miocene. They contain several species of beavers. Amongst them are S.barbouri, S.complexus, S.depereti, S.fossor, S.gradatus, and S.hesperus. Their various species are found all the way from the eastern end of the Iberian peninsula to southern Japan. S.depereti has been found in northwest Germany.
Daphoenus is an extinct genus of amphicyonids. Daphoenus inhabited North America from the Late Eocene to the Middle Miocene, 37.2—16.0 Mya, existing for approximately 21 million years.
The zygomasseteric system in rodents is the anatomical arrangement of the masseter muscle of the jaw and the zygomatic arch of the skull. The anteroposterior or propalinal (front-to-back) motion of the rodent jaw is enabled by an extension of the zygomatic arch and the division of the masseter into a superficial, lateral and medial muscle. The four main types are described as protrogomorphous, sciuromorphous, hystricomorphous, and myomorphous.
Parictis is an extinct arctoid belonging to the family Subparictidae.
Castor californicus is an extinct species of beaver that lived in western North America from the end of the Miocene to the early Pleistocene. Castor californicus was first discovered in Kettleman Hills in California, United States. The species was similar to but larger than the extant North American beaver, C. canadensis.
Anchitheriomys is an extinct member of the beaver family, Castoridae. It inhabited North America and Eurasia during the middle Miocene. The name of the genus comes from Anchitherium, an extinct genus of horses, and the Greek word for mouse, μῦς (mys), thus meaning "Anchitherium's mouse", because the fossils of both genera usually co-occur.
Propalaeocastor is a poorly known extinct genus of beavers from the early Oligocene of Europe and Asia. Recently described material of a new species of Propalaeocastor, P. irtyshensis, indicates the genus is probably the earliest known member of the subfamily Castorinae, which includes all castorids more closely related to living beavers than to the extinct giant beaver. Previously, Propalaeocastor had been allied with more basal beavers such as Agnotocastor and Anchitheriomys. According to Wu and coauthors, Propalaeocastor, specifically P. butselensis, is the likely ancestor of the better-known Eurasian castorine Steneofiber.
Migmacastor is an extinct member of the beaver family, Castoridae, known from a single species, Migmacastor procumbodens. Only a single specimen has been reported, a skull from the late Oligocene or early Miocene of Nebraska. Features of the incisor teeth of Migmacastor indicate they were used to dig. Other extinct beavers, including the better-known Palaeocastor, were also fossorial (digging), but Migmacastor may have become a burrower independently.
Cormocyon is an extinct genus of borophagine canid native to North America. It lived from the Oligocene to the Early Miocene, 30.8—20.6 Mya, existing for about 10.2 million years. It is regarded as a primitive, transitional member of the Borophagini tribe.
Trogontherium is an extinct genus of Eurasian giant beavers that lived from the Late Pliocene to the Late Pleistocene. Fossils of Trogontherium have been found across northern Eurasia, from Western Europe to China and Siberia.
Brachyhyops is an extinct genus of entelodont artiodactyl mammal that lived during the Eocene Epoch of western North America and southeastern Asia. The first fossil remains of Brachyhyops are recorded from the late Eocene deposits of Beaver Divide in central Wyoming and discovered by paleontology crews from the Carnegie Museum of Natural History during the early 20th century. The type species, Brachyhyops wyomingensis, is based on a single skull and was named by E.H. Colbert in 1937, but was not officially described until 1938. During the latter half of the 20th century, additional specimens from North America have been recorded from Saskatchewan and as far south as Texas, indicating that Brachyhyops had a broad distribution and was well-dispersed throughout western North America.
Diegoaelurus is an extinct genus of placental mammals from extinct the subfamily Machaeroidinae within extinct family Oxyaenidae. This genus contains only one species Diegoaelurus vanvalkenburghae, which was found in the Santiago Formation in California. This mammal lived during the Uintan stage of the Middle Eocene Epoch around 46.2 to 39.7 million years ago.
