Alienopteridae Temporal range: | |
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Generic reconstruction of an alienopterid adult | |
Aethiocarenus , an alienopterid nymph | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Superfamily: | † Umenocoleoidea |
Family: | † Alienopteridae Bai et al, 2016 |
Type genus | |
† Alienopterus Bai et al, 2016 | |
Genera | |
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Synonyms | |
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Alienopteridae is an extinct family of dictyopterans, known from the Early Cretaceous to the Eocene epoch. They are noted for their unusual combination of features not found in other dictyopterans.
Alienopterid nymphs vary in morphology, some are superficially ant-like, while others are more robust. In adult alienopterids, the first set of wings are reduced into shortened sclerotised scale-like structures. The compound eyes are large, and the head generally triangular shaped and able to rotate. [1]
It was originally assigned to its own order Alienoptera by Bai et al., 2016. It was reassigned to the dictyopteran superfamily Umenocoleoidea as sister family to the beetle-like Umenocoleidae by Vršanský et al. (2018), [2] and a more recent analysis similarly places Alienopteridae and Umenocoleidae as sister taxa within Dictyoptera, but placing both lineages outside of Blattodea. [3] A 2021 study revived the order Alienoptera for the clade containing Alienopteridae and Umenocoleidae, with a cladistic analysis placing Alienoptera as the sister clade to Mantodea (praying mantises). [1]
The majority of the alienopterid genera are known from the mid Cretaceous (latest Albian-earliest Cenomanian ~ 100 million years ago) Burmese amber found in Myanmar; [4] [5] [6] [2] [7] [8] though an additional two genera ( Apiblatta and Vcelesvab ) are from the late Aptian Crato Formation (Brazil), and the two youngest genera ( Chimaeroblattina and Grant ) are from the middle Eocene Green River Formation (Colorado, United States). [1]
In a 2018 paper, some alienopterid adults were suggested as mimics of various hymenopterans, including bees and ants. [2] However this was mostly rejected in another study as lacking evidence. [9] A 2021 study alternatively suggested that some alienopterid nymphs functioned as ant mimics, based on morphological features that closely resembled contemporary sphecomyrmine ants, and suggested that the adult genus Teyia was also a wasp mimic, though mimicry in other adult alienopterids was rejected. One alienopterid nymph was found laden with gymnosperm pollen (probably from a cycad or a bennettitalean), suggesting that it consumed pollen as part of its diet, and that it also acted as a pollinator. [1] Alienopterid adults have been suggested to be predators adapted to moving through dense foliage, [9] with the small sclerotised forewings being an adaptation for flight capability. [1]
An undescribed species is also known from the Turonian aged Orapa kimberlite pipe sediments in Botswana. [2] Luo, Xu & Jarzembowski (2020) transferred Alienopterix and Vzrkadlenies, originally described as alienopterids, to the family Cratovitismidae. [10] A 2021 study later considered Alienopterix an umenocoleid instead. [1]
The insect order Zoraptera, commonly known as angel insects, contains small and soft bodied insects with two forms: winged with wings sheddable as in termites, dark and with eyes (compound) and ocelli (simple); or wingless, pale and without eyes or ocelli. They have a characteristic nine-segmented beaded (moniliform) antenna. They have mouthparts adapted for chewing and are mostly found under bark, in dry wood or in leaf litter.
Dictyoptera is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea and the order Mantodea (mantises). All modern Dictyoptera have short ovipositors and typically lay oothecae. The oldest fossils of Dictyoptera from the Late Carboniferous, referred to as "roachoids" have long ovipositors and did not lay oothecae. The oldest modern oothecae-laying dictyopterans date to the Late Triassic.
Corydiidae, previously known as Polyphagidae, is a family of the order Blattodea (cockroaches). Many are known as sand cockroaches. The family is divided into five subfamilies, comprising some 40 genera. One prominent species is the desert cockroach, Arenivaga investigata.
Blattodea is an order of insects that contains cockroaches and termites. Formerly, termites were considered a separate order, Isoptera, but genetic and molecular evidence suggests they evolved from within the cockroach lineage, cladistically making them cockroaches as well. The Blattodea and the mantis are now all considered part of the superorder Dictyoptera. Blattodea includes approximately 4,400 species of cockroach in almost 500 genera, and about 3,000 species of termite in around 300 genera.
Corydioidea is a superfamily of insects in the order Blattodea containing the cockroaches and termites. It contains two extant families, Corydiidae and Nocticolidae, comprising about fifty genera and two hundred and fifty species, along with the extinct family Liberiblattinidae. Members of this superfamily are found worldwide, mostly in hot, arid habitats.
Alienopterus brachyelytrus is an extinct insect described from a 99 million year old fossil found in Burmese amber from the Hukawng Valley of Myanmar. It was the first known member of the order Alienoptera until 2018, when the second and third members of the order, Caputoraptor elegans, and Alienopterella stigmatica were described. A. brachyelytrus has characters that are shared with cockroaches and mantids and is thought to represent either the sister taxon, or an ancestor to mantids.
Aethiocarenus is an extinct genus of insects which has a single species Aethiocarenus burmanicus described from a 98.79 ±0.62 million year old fossil found in Burmese amber from the Hukawng Valley of Myanmar. The insect is unusual due to the vertex of the triangular head being attached to the pronotum as opposed to the hypotenuse. When first described Aethiocarenus was placed as the sole member of the family Aethiocarenidae and order Aethiocarenodea. However, Aethiocarenus was later considered to be a nymph of Alienopterus. Vršanský et al. (2018) considered Aethiocarenus to be an alienopterid nymph, but considered it distinct from other members of this group and deserving a separate genus rank.
Caputoraptor is an extinct genus of insect from the Cenomanian aged Burmese amber containing two species: the type species Caputoraptor elegans, as well as Caputoraptor vidit. It is part of the extinct order Alienoptera. C. elegans is notable for the presence of a scissor like mechanism consisting of a straight edge on the back of the head and corresponding serrated edges on the first thoracic segment, these were initially suggested to be used by the female to grasp the male during mating, but the structure is not sexually dimorphic, so a use to hold prey was subsequently suggested. Its morphology suggests a predatory habit inhabiting shrubs and trees. In 2020 a Caputoraptor elegans nymph was described that was in the process of being predated upon by a Ceratomyrmex hell ant.
2018 in paleoentomology is a list of new fossil insect taxa that were described during the year 2018, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
2019 in paleoentomology is a list of new fossil insect taxa that were described during the year 2019, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
2017 in paleoentomology is a list of new fossil insect taxa that were described during the year 2017, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
2020 in paleoentomology is a list of new fossil insect taxa that were described during the year 2020, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
2015 in paleoentomology is a list of new fossil insect taxa that were described during the year 2016, as well as other significant discoveries and events related to paleoentomology that were scheduled to occur during the year.
Burmese amber is fossil resin dating to the early Late Cretaceous Cenomanian age recovered from deposits in the Hukawng Valley of northern Myanmar. It is known for being one of the most diverse Cretaceous age amber paleobiotas, containing rich arthropod fossils, along with uncommon vertebrate fossils and even rare marine inclusions. A mostly complete list of all taxa described up until 2018 can be found in Ross 2018; its supplement Ross 2019b covers most of 2019.
Umenocoleidae is an extinct family of dictyopteran insects known from the Cretaceous. They are considered to be closely related to the Alienopteridae. They were originally considered to be beetles due to their beetle-like morphology, with sclerotised elytra-like forewings. This was probably an adaptation for living under bark and in other tight spaces.
Liberiblattinidae is an extinct family of cockroaches known from the Jurassic to Cretaceous. Some taxa, like Cryptoblatta and Hydrokhoohydra, are suggested to be semiaquatic. Spongistoma is suggested to be a nectarivore due to its unique sucking/sponging "proboscis" mouthparts. Some authors have suggested that the family is ancestral to Mantodea.
Elcanidae are an extinct family of Mesozoic and early Cenozoic orthopterans. Members of the family are distinguished by the presence of spurs on the distal part of the metatibia, unique among orthopterans, these have been suggested to have been used for controlling gliding, swimming aids, or for jumping on water. The group combines characteristics from both major groups of orthopterans, with long antennae and nymphal morphology similar to Ensifera, but with wing venation and adult morphology more similar to Caelifera. Elcanidae is part of Elcanoidea, which is thought to have diverged from living orthopterans by the beginning of the Permian, around 300 million years ago. The family also includes Permelcanidae, known from the Early-Late Permian. The relationship of Elcanoidea to Ensifera and Caelifera is currently unresolved. Elcanids are known from the Late Triassic to Paleocene of Eurasia, North and South America. Some members of the group exhibited aposematic coloration. They are thought to have been herbivorous.
Phygasia is a genus of flea beetles in the family Chrysomelidae. There are around 70 described species in Phygasia, found in the Palearctic, Indomalaya, and tropical Africa.
This paleoentomology list records new fossil insect taxa that are to be described during the year 2022, as well as notes other significant paleoentomology discoveries and events which occurred during that year.
This paleoentomology list records new fossil insect taxa that were described during the year 2014, as well as notes other significant paleoentomology discoveries and events which occurred during that year.