Madagascan warblers | |
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The long-billed bernieria (Bernieria madagascariensis) was formerly placed in the Pycnonotidae | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Order: | Passeriformes |
Superfamily: | Sylvioidea |
Family: | Bernieridae Cibois, David, Gregory & Pasquet, 2010 [1] |
Genera | |
see text |
The tetrakas and allies are a newly validated family of songbirds. They were formally named Bernieridae in 2010. The family currently consists of eleven species (in eight genera) of small forest birds. These birds are all endemic to Madagascar.
In 1934, the monophyly of this group was proposed by Finn Salomonsen but the traditional assignments of these birds were maintained, mistaken by their convergent evolution and the lack of dedicated research. The families to which the Malagasy warblers were formerly assigned—Pycnonotidae (bulbuls) and even more so Timaliidae (Old World babblers) and the Old World warbler—were used as "wastebin taxa", uniting unrelated lineages that were somewhat similar ecologically and morphologically.
It was not until the analysis of mtDNA cytochrome b and 16S rRNA [2] [3] as well as nDNA RAG-1 and RAG-2 exon sequence data, [4] that the long-proposed grouping was accepted. [5]
The family contains 11 species divided into 8 genera. [5]
Image | Genus | Species |
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Oxylabes Sharpe, 1870 |
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Bernieria Pucheran, 1855 |
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Cryptosylvicola Goodman, Langrand & Whitney, 1996 |
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Hartertula Stresemann, 1925 |
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Thamnornis Milne-Edwards & Grandidier, 1882 |
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Xanthomixis Sharpe, 1881 |
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Crossleyia Hartlaub, 1877 |
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Randia Delacour & Berlioz, 1931 |
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Several of these species are very poorly known and were described by science only very recently. Appert's tetraka was only described in 1972 and the cryptic warbler in 1996. The Appert's tetraka, along with the dusky tetraka are threatened by habitat loss, and are listed as vulnerable.
Most members of this family live in the humid rainforests in the east of Madagascar, though a few species are found in the drier southwest of the island. They feed on insects and will form mixed-species feeding flocks of up to six species while foraging.
A passerine is any bird of the order Passeriformes, which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their toes, which facilitates perching.
The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical Asia to Indonesia, and north as far as Japan. A few insular species occur on the tropical islands of the Indian Ocean. There are 166 species in 32 genera. While different species are found in a wide range of habitats, the African species are predominantly found in rainforest, whereas Asian bulbuls are predominantly found in more open areas.
Old World warblers are a large group of birds formerly grouped together in the bird family Sylviidae. They are not closely related to the New World warblers. The family held over 400 species in over 70 genera, and were the source of much taxonomic confusion. Two families were split out initially, the cisticolas into Cisticolidae and the kinglets into Regulidae. In the past ten years they have been the subject of much research and many species are now placed into other families, including the Acrocephalidae, Cettiidae, Phylloscopidae, and Megaluridae. In addition some species have been moved into existing families or have not yet had their placement fully resolved. A smaller number of warblers, together with some babblers formerly placed in the family Timaliidae and the parrotbills, are retained in a much smaller family Sylviidae.
The typical warblers are small birds belonging to the genus Sylvia in the "Old World warbler" family Sylviidae.
The Old World babblers or Timaliidae are a family of mostly Old World passerine birds. They are rather diverse in size and coloration, but are characterised by soft fluffy plumage. These are birds of tropical areas, with the greatest variety in Southeast Asia and the Indian subcontinent. The timaliids are one of two unrelated groups of birds known as babblers, the other being the Australasian babblers of the family Pomatostomidae.
The parrotbills are a family, Paradoxornithidae, of passerine birds that are primarily native to East and Southeast Asia, though feral populations exist elsewhere. They are generally small, long-tailed birds that inhabit reedbeds and similar habitats. They feed mainly on seeds, e.g. of grasses, to which their bill, as the name implies, is well-adapted. Living in tropical to southern temperate climates, they are usually non-migratory.
The family Vangidae comprises a group of often shrike-like medium-sized birds distributed from Asia to Africa, including the vangas of Madagascar to which the family owes its name. Many species in this family were previously classified elsewhere in other families. Recent molecular techniques made it possible to assign these species to Vangidae, thereby solving several taxonomic enigmas. The family contains 40 species divided into 21 genera.
The family Cisticolidae is a group of about 160 warblers, small passerine birds found mainly in warmer southern regions of the Old World. They were formerly included within the Old World warbler family Sylviidae.
The white-eyes are a family, Zosteropidae, of small passerine birds native to tropical, subtropical and temperate Sub-Saharan Africa, southern and eastern Asia, and Australasia. White-eyes inhabit most tropical islands in the Indian Ocean, the western Pacific Ocean, and the Gulf of Guinea. Discounting some widespread members of the genus Zosterops, most species are endemic to single islands or archipelagos. The silvereye, Zosterops lateralis, naturally colonised New Zealand, where it is known as the "wax-eye" or tauhou ("stranger"), from 1855. The silvereye has also been introduced to the Society Islands in French Polynesia, while the Japanese white-eye has been introduced to Hawaii.
The wrentit is a small bird that lives in chaparral, oak woodlands, and bushland on the western coast of North America. It is the only species in the genus Chamaea.
The greenbuls are a group of birds within the bulbul family Pycnonotidae, found only within Africa. They are all largely drab olive-green above, and paler below, with few distinguishing features.
Appert's tetraka formerly known as Appert's greenbul is a small passerine bird endemic to the south-west of Madagascar. The species was only described in 1972, and has been the subject of considerable taxonomic confusion. It was initially placed in the greenbul genus Phyllastrephus, and later with the Old World warblers in the genus Bernieria. Recent research indicates it is part of an endemic Malagasy radiation currently known as the Malagasy warblers.
The wedge-tailed jery is a small bird endemic to the east of Madagascar. The species has been the cause of some taxonomic confusion, it was originally placed with the jeries in the genus Neomixis (Cisticolidae) before being placed in its own monotypic genus Hartertula, but still considered close to Neomixis. Recent research indicates it is part of an endemic Malagasy radiation currently known as the Malagasy warblers.
The long-billed bernieria, formerly known as long-billed greenbul and sometimes as common tetraka or long-billed tetraka, is a songbird species endemic to Madagascar. It is the only species placed in the genus Bernieria. Its natural habitat is subtropical or tropical moist lowland forests.
The white-throated oxylabes is a species of passerine bird that is endemic to Madagascar. It is the only species placed in the genus Oxylabes. Formerly considered as a member of the Old World warbler family Sylviidae, it has been moved to the family Bernieridae — the Malagasy warblers. Its natural habitat is subtropical or tropical moist lowland forests.
Locustellidae is a newly recognized family of small insectivorous songbirds ("warblers"), formerly placed in the Old World warbler "wastebin" family. It contains the grass warblers, grassbirds, and the Bradypterus "bush warblers". These birds occur mainly in Eurasia, Africa, and the Australian region. The family name is sometimes given as Megaluridae, but Locustellidae has priority.
Neomixis is a genus of small forest birds that are endemic to Madagascar.
Pnoepyga is a genus of passerines endemic to southern and southeastern Asia. Its members are known as cupwings or wren-babblers. The genus contains four species. The genus has long been placed in the babbler family Timaliidae. A 2009 study of the DNA of the families Timaliidae and the Old World warblers (Sylviidae) found no support for the placement of the genus in either family, prompting the authors to erect a new monogeneric family, the Pnoepygidae.
The Beijing babbler, also known as the white-browed Chinese warbler, Chinese hill warbler, or Chinese bush-dweller, is a species of bird in the genus Rhopophilus. It is now thought to be a close relative of the parrotbills and is placed in the family Paradoxornithidae; previously, it was placed in the families Cisticolidae, Timaliidae or Sylviidae. It is found in northern China and North Korea, and formerly occurred in South Korea. The species was first described by Robert Swinhoe in 1868.
Sylvioidea is a superfamily of passerine birds, one of at least three major clades within the Passerida along with the Muscicapoidea and Passeroidea. It contains about 1300 species including the Old World warblers, Old World babblers, swallows, larks and bulbuls. Members of the clade are found worldwide, but fewer species are present in the Americas.