Bolbolenellus

Bolbolenellus ; Temporal range: Toyonian (Upper Olenellus-zone) 516–513 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Trilobita
Order:	Redlichiida
Suborder:	Olenellina
Superfamily:	Olenelloidea
Family:	Biceratopsidae
Subfamily:	Biceratopsinae
Genus:	Bolbolenellus; Palmer & Repina, 1993
Species
	B. euryparia(Palmer, 1979) (Type), synonyms Olenellus euryparia, O. fremonti pro parte ; B. altifrons(Fritz, 1972) , synonym Olenellus altifrons; B. brevispinusPalmer, 1998; B. groenlandicus(Poulsen, 1927), synonyms Olenellus groenlandicus, O. kentensis, Bristolia groenlandicus, Bristolia kentensis; B. hermani(Kindle & Tasch, 1948), synonyms Olenellus hermani, Esmeraldina hermani; B. sphaerulosus(Fritz, 1991), synonyms Olenellus sphaerulosus, O. bufrontis;

Last updated July 18, 2024

Bolbolenellus is an extinct genus of trilobites, fossil marine arthropods, with five species attributed to it currently. It can be easily distinguished from all other trilobites by the combination of the absence of dorsal sutures in the head shield like all Olenellina (which in all other trilobite suborders assist in moulding by splitting open), and a distinctly bulbous frontal lobe (L4) of the raised axial area in the head (or cephalon) called glabella. The species lived at the end of the Lower Cambrian.

Etymology

Bolbolenellus is the combination of Greek βολβός (bolbos, “plant with round swelling on underground stem”), and Olenellus , the rather distantly related genus to which all of the species were previously assigned. This refers to the bulb-like swelling of the frontal lobe of the glabella. The names of the species have the following derivations.

B. altifrons from the Latin altus (“high”) and from Latin frons (“forehead”), to express the species has a bulbous frontal glabellar lobe.
B. brevispinus from brevis the Latin word for "short" and spinus the Latin word for "spine".
B. groenlandicus is named for Greenland, where the species occurs.
B. sphaerulosus from the Latin sphaerula (small ball).

Description

As with most early trilobites, Bolbolenellus has an only thinly calcified exoskeleton. Although most of the body is rather flat, the frontal lobe (L4) of the central area of the cephalon (or glabella) is prominent, bulbous (or subglobular), and may overlap the border at the front in dorsal view. It also shares crescent-shaped eye ridges, but these rise above the exoskeleton. As part of the Olenellina suborder, Bolbolenellus lacks dorsal sutures. Like all other members of the Olenelloidea superfamily, the eye-ridges spring from the back of the frontal lobe (L4) of the glabella. The posterior margin of cephalon is nearly straight or angles to the front (anterolaterally) on the outside of the intergenal spine or intergenal angle. The short genal spine protrudes opposite or posterior to the most backward side lobe of the glabella (or L1). There may be an intergenal spine. The frontal lobe of the glabella touches the border of the cephalon (or the preglabellar field is absent). The most backward tips of eye ridges (or ocular lobes) are approximately opposite occipital furrow. Although the third thorax) segment (or T3, this is counted from the headshield to the tailshield) has strongly enlarged side lobes (or pleurae), the spine on their end is not much enlarged compared to the spines of neighboring segments. The 14th segment carries a backward pointing axial spine that is approximately as long as the thorax axis. There seems to be no opistothorax.^[1]^[2]

Differences with some other Biceratopsinae

Most other Biceratopsinae, i.e. Biceratops nevadensis , Emigrantia and Peachella and to a lesser extend Eopeachella have effaced cephalic features. Biceratops lacks genal spines. In Emigrantia the genal spines are longer than the cephalon and attach halfway down its side (or lateral margin). Peachella has club-like genal spines.^[3] In Nephrolenellus the eye ridges extend outwards, the second pair of side lobes of the glabella are very narrow, the genal spines attach further to the front of the cephalon, T3 carries massive pleural spines, the axial spine on T14 is absent, and there is a prominent opistothorax.^[2]

Relations with other Biceratopsinae

The closest relative of the six species of Bolbolenellus is the genus Nephrolenellus. The other genera of the Biceratopsinae (Biceratops, Emigrantia, Eopeachella, Peachella, and Olenelloides) constitute the sister group.^[1]

Distribution

B. euryparia is present in the Upper Olenellus-zone of California (lower half of the B. euryparia-subzone of the Carrara Formation, Eagle Mountain, White-Inyo Mountains, and the middle part of the Latham Shale, East slope of hill 1440 in the southern end of the Marble Mountains, San Bernardino County).^[1]
B. altifrons occurs in the Middle Olenellus-zone of the Northwest Territories (type section of the type section of the Sekwi Formation, 0.2 miles Southeast of June Lake, Mackenzie Mountains).^[1]
B. brevispinus has been collected in the Upper Olenellus-zone of Nevada (near Oak Spring Summit, Pioche Formation, Lincoln County).^[2]
B. groenlandicus is found in Northwestern Greenland, questionably from the Upper Olenellus-zone (Wulff River Formation, Marshall Bugt, Inglefield Land).^[1]
B. hermani occurs in the Upper Olenellus-zone of Vermont (Monkton Quartzite, near Clay and Red Rock Points on Lake Champlain, and Parker Slate, 2 miles west of Georgia Centre), of California (White|Inyo Mountain region)^[1]
B. sphaerulosus has been collected in the Olenellus-zone of the Yukon Territory, Canada (Unit 6 of the Upper Illtyd Formation, Wernecke Mountains).^[1]^[4]

Habitat

Bolbolenellus was probably a marine bottom dweller, like all Olenellina.

Related Research Articles

<span class="mw-page-title-main">Redlichiida</span> Extinct order of trilobites

Redlichiida is an order of trilobites, a group of extinct marine arthropods. Species assigned to the order Redlichiida are among the first trilobites to appear in the fossil record, about halfway during the Lower Cambrian. Due to the difficulty to relate sediments in different areas, there remains some discussion, but among the earliest are Fallotaspis, and Lemdadella, both belonging to this order. The first representatives of the orders Corynexochida and Ptychopariida also appear very early on and may prove to be even earlier than any redlichiid species. In terms of anatomical comparison, the earliest redlichiid species are probably ancestral to all other trilobite orders and share many primitive characters. The last redlichiid trilobites died out before the end of the Middle Cambrian.

Olenellus is an extinct genus of redlichiid trilobites, with species of average size. It lived during the Botomian and Toyonian stages (Olenellus-zone), 522 to 510 million years ago, in what is currently North-America, part of the palaeocontinent Laurentia.

<span class="mw-page-title-main">Emuellidae</span> Extinct family of trilobites

Emuellidae are a small family of trilobites, a group of extinct marine arthropods, that lived during the late Lower Cambrian of the East Gondwana supercontinent, in what are today South-Australia and Antarctica.

Olenelloides armatus is an extinct, small sized olenelloid redlichiid trilobite arthropod. It lived during the later part of the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period. The most conspicuous feature is the hexagonal head shield that carries 6 ray-like spines..

Wanneria is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the later part of the Botomian stage, which lasted from approximately 524 to 518.5 million years ago. This faunal stage was part of the Cambrian Period. Wanneria walcottana is the only known species in this genus.

Elliptocephala is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived from the later part of the Tommotian to the upper Botomian. Elliptocephala can easily be confused with Ellipsocephalus, a trilobite genus of the Ptychopariida order.

Fremontella is an extinct genus from a well-known class of fossil marine arthropods, the trilobites. It lived during the part of the Toyonian stage. This faunal stage was part of the Cambrian Period. Fremontella shares with the other genera of the Bristoliinae subfamily, Lochmanolenellus and Bristolia conspicuous and long curved spines on the headshield. These reach back equal to 4-5 thorax segments. The furrows that separate border, eye ridges, glabella and its lobes are distinct. The area outside of the axis of the third segment of the thorax is enlarged, and carries large trailing spine on each side.

Archaeaspis is an extinct genus of redlichiid trilobites. It lived during the late Atdabanian stage, which lasted from 521 to 514 million years ago during the early part of the Cambrian Period. The first specimens were first found in Siberia, where they were originally thought to have been endemic. Later, though, more specimen were found in the Inyo Mountains in Southern California. An Ediacarian proarticulatan was given the same name, Archaeaspis Ivantsov, 2001. This is however a junior homonym. The name Archaeaspis Repina in Khomentovskii and Repina, 1965, for the trilobite has priority. The new valid name for the proarticulate is Archaeaspinus Ivantsov, 2007.

Nephrolenellus is an extinct genus of trilobite, fossil marine arthropods, of relatively small size. Currently two species are attributed to it. Nephrolenellus lived at the end of the Lower Cambrian. Species are known from the Great Basin of California, Nevada and Arizona, with one specimen from Canada.

Mesonacis is an extinct genus of trilobite that lived during the Botomian, found in North-America, and the United Kingdom. Some of the species now regarded part of Mesonacis, have previously been assigned to Angustolenellus or Olenellus (Angustolenellus). Angustolenellus is now regarded a junior synonym of Mesonacis.

<span class="mw-page-title-main">Holmiidae</span> Family of trilobites

Holmiidae is a family of trilobites, that lived during the Lower Cambrian (Atdabanian). The Holmiidae is a diverse family of eight genera containing at least 17 species. It includes some of the earliest trilobites of Baltica. Holmiidae occur throughout Baltica and Western Laurentia, and also in Morocco.

Peachella is an extinct genus of trilobites, fossil marine arthropods, with species of average size. It lived during the Toyonian stage, 516 to 513 million years ago, in what is today the southwestern United States. It can easily be distinguished from other trilobites by its club-like genal spines.

Eopeachella is an extinct genus of trilobites, fossil marine arthropods, with species of probably small size, entire, articulate specimens have not yet been found. It lived during the Toyonian stage, in what is today the South-Western United States. Eopeachella angustispina is the only known species in this genus.

The Biceratopsinae is an extinct subfamily of redlichiid trilobites within the family Biceratopsidae, with species of small to average size. Species belonging to this subfamily lived during the Toyonian stage, 516-513 million years ago, in the former continent of Laurentia, including what are today the South-Western United States and Canada.

Biceratops is an extinct genus of olenelloid redlichiid trilobites, of average size, with the largest specimen 8 centimetres or 3.1 inches long, not including the huge pleural spines of the 3rd segment of the thorax. It lived during the Toyonian stage, in what is today the South-Western United States. Biceratops can easily be distinguished from other members of Biceratopsidae by the absence of genal spines, in combination with effaced features of the raised axial area of the head shield, that is bordering the two horn-like projections that carry the eyes. Biceratops nevadensis is the only known species in this genus.

Mesolenellus is an extinct genus of trilobites that lived during the lower Cambrian (Botomian), found in Greenland and Spitsbergen.

The Bristoliinae is an extinct subfamily of trilobites, fossil marine arthropods, with species of small to average size. Species belonging to this subfamily lived during the Botomian and Toyonian stage (Olenellus-zone), 522-513 million years ago, in the former continent of Laurentia, including what are today Mexico, the Appalachian Mountains and the south-western United States, and Canada.

Lochmanolenellus is an extinct genus of trilobites, fossil marine arthropods, with one small species, L. mexicana. It lived during the Botomian stage (Olenellus-zone), 522–513 million years ago, in the South-West of the former continent of Laurentia, in what are today Mexico, and the South-Western United States.

Bristolia is an extinct genus of trilobite, fossil marine arthropods, with eight or more small to average size species. It is common in and limited to the Lower Cambrian shelf deposits across the southwestern US, which constitutes part of the former paleocontinent of Laurentia.

Emigrantia is an extinct genus of trilobites, fossil marine arthropods, of small to average size. It lived during the Toyonian stage, in what is today the South-Western United States. Emigrantia can easily be distinguished from other trilobites by the sturdy but not inflated genal spines, that are attached at midlength of the cephalon, in combination with effaced features of the raised axial area of the head shield.

References

1 2 3 4 5 6 7 8 9 10 11 12 13 Lieberman, B.S. (1999). "Systematic Revision of the Olenelloidea (Trilobita, Cambrian)" (PDF). Bulletin of the Peabody Museum of Natural History. 45.
1 2 3 4 Palmer, A.R. (1998). "Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada". Journal of Paleontology. 72 (4): 650–672. Bibcode:1998JPal...72..650P. doi:10.1017/S0022336000040373. JSTOR 1306693. S2CID 131861504.
↑ Mark Webster (2011). Hollingsworth, J.S.; Sundberg, F.A.; Foster, J.R. (eds.). "Trilobite Biostratigraphy and Sequence Stratigraphy of the Upper Dyeran (traditional Laurentian "Lower Cambrian) in the southern Great Basis, USA". Museum of Northern Arizona Bulletin. 67– via Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada.
↑ Fritz, W.H. (1991), "Lower Cambrian Trilobites from the Illtyd Formation, Wernecke Mountains, Yukon Territoriy", Bulletin of the Geological Survey of Canada, 409: 1–77

External links

This page is based on this Wikipedia article
Text is available under the CC BY-SA 4.0 license; additional terms may apply.
Images, videos and audio are available under their respective licenses.

[Lieberman_1999-1] 1 2 3 4 5 6 7 8 9 10 11 12 13 Lieberman, B.S. (1999). "Systematic Revision of the Olenelloidea (Trilobita, Cambrian)" (PDF). Bulletin of the Peabody Museum of Natural History. 45.

[Palmer-2] 1 2 3 4 Palmer, A.R. (1998). "Terminal Early Cambrian Extinction of the Olenellina: Documentation from the Pioche Formation, Nevada". Journal of Paleontology. 72 (4): 650–672. Bibcode:1998JPal...72..650P. doi:10.1017/S0022336000040373. JSTOR 1306693. S2CID 131861504.

[3] Mark Webster (2011). Hollingsworth, J.S.; Sundberg, F.A.; Foster, J.R. (eds.). "Trilobite Biostratigraphy and Sequence Stratigraphy of the Upper Dyeran (traditional Laurentian "Lower Cambrian) in the southern Great Basis, USA". Museum of Northern Arizona Bulletin. 67– via Cambrian Stratigraphy and Paleontology of Northern Arizona and Southern Nevada.

[4] Fritz, W.H. (1991), "Lower Cambrian Trilobites from the Illtyd Formation, Wernecke Mountains, Yukon Territoriy", Bulletin of the Geological Survey of Canada, 409: 1–77

[1]

[2]

[3]

[4]