Cimolichthys Temporal range: | |
---|---|
C. nepaholica mount, University of Michigan Museum of Natural History | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Aulopiformes |
Superfamily: | † Cimolichthyoidea |
Family: | † Cimolichthyidae Goody, 1969 |
Genus: | † Cimolichthys Leidy, 1857 |
Type species | |
†Cimolichthys levesiensis Leidy, 1857 | |
Species | |
| |
Synonyms | |
|
Cimolichthys (Greek for "chalk fish") is an extinct genus of large predatory marine aulopiform ray-finned fish known worldwide from the Late Cretaceous. It is the only member of the family Cimolichthyidae. [2] [3]
Cimolichthys was a large-sized nektonic aulopiform fish, making it related to modern lancetfish and lizardfish. Within the Aulopiformes, it is generally considered a member of the Enchodontoidei, a dominant group of predatory nektonic fish throughout much of the Cretaceous; [3] [4] however, some other treatments instead place it outside the Enchodontoidei and in a basal position as sister to the waryfishes, a small family of extant deep-sea aulopiformes. [5] Yet other studies have instead placed it as a sister to the extant daggertooths. [6]
The following species are known: [7]
C. anceps, C. gladiolus, C. semianceps, C. contracta, C. merillii, and C. sulcatus, all described by Cope (1872), are now considered synonymous with C. nepaholica. [3] [12] Indeterminate remains are known from the Maastrichtian of Niger & Brazil, and the Turonian of the Czech Republic. [7] [13]
Although the closest living relatives of Cimolichthys are lancetfish and lizardfish, the living animals would have resembled very large pikes. They could grow up to 1.5 to 2.0 metres (4.9 to 6.6 ft) meters long. Their bodies were covered by large, heavy scutes. Typical of this species are narrow lower jaws with several series of teeth. Remains of undigested fishes or squids have been found in collected specimens. Particularly well-preserved specimens are known from the Niobrara Formation of Kansas. [14] Their barbed teeth are distinctive, and can identify the presence of this genus even in formations where articulated remains are not otherwise known. [11]
Aulopiformes is a diverse order of marine ray-finned fish consisting of some 15 extant and several prehistoric families with about 45 genera and over 230 species. The common names grinners, lizardfishes and allies, or aulopiforms are sometimes used for this group. The scientific name means "Aulopus-shaped", from Aulopus + the standard fish order suffix "-formes". It ultimately derives from Ancient Greek aulós + Latin forma, the former in reference to the elongated shape of many aulopiforms.
The Myctophiformes are an order of ray-finned fishes consisting of two families of deep-sea marine fish, most notably the highly abundant lanternfishes (Myctophidae). The blackchins (Neoscopelidae) contain six species in three genera, while the bulk of the family belongs to the Myctophidae, with over 30 genera and some 252 species.
Enchodus is an extinct genus of aulopiform ray-finned fish related to lancetfish and lizardfish. Species of Enchodus flourished during the Late Cretaceous, and there is some evidence that they may have survived to the Paleocene or Eocene; however, this may just represent reworked Cretaceous material.
Apateodus is a genus of prehistoric marine ray-finned fish which was described by Woodward in 1901. It was a relative of modern lizardfish and lancetfish in the order Aulopiformes, and one of a number of prominent nektonic aulopiforms of Cretaceous marine ecosystems.
Apateopholis is an extinct genus of prehistoric marine ray-finned fish. It is the only member of the family Apateopholidae and contains a single species, A. laniatus, from the Cenomanian of Lebanon. It was a relative of modern lizardfish and lancetfish in the order Aulopiformes. At least one study has found it to be a potential sister genus to the enigmatic Yabrudichthys of the West Bank.
Cyranichthys is an extinct genus of marine ray-finned fish known from the Late Cretaceous of central Africa and western Europe. It was a member of Dercetidae, a group of elongated aulopiforms.
Dercetis is a genus of prehistoric marine ray-finned fish. It is the type genus of the family Dercetidae, a group of slender, elongate aulopiforms, which were related to modern lizardfish and grinners. It is known from the Late Cretaceous of Europe, the Middle East, and western North America.
Dercetoides is a genus of prehistoric marine ray-finned fish from the Late Cretaceous period. It contains a single species, D. venator, known from the early Cenomanian-aged Amminadav Formation of the West Bank, Palestine. It was a member of the Dercetidae, a group of elongated aulopiforms that were related to modern lancetfish and lizardfish.
Prionolepis is a genus of prehistoric ray-finned fish belonging to the order Alepisauriformes.
Bananogmius is an extinct genus of marine ray-finned fish that was found in what is now North America and Europe during the Late Cretaceous, from the Cenomanian to the Santonian. It lived in the Western Interior Seaway, which split North America in two during the Late Cretaceous, as well as the proto-North Sea of Europe.
Aulolepis is an extinct genus of prehistoric marine ray-finned fish that lived from the middle Cenomanian to the late Turonian. It contains a single species, A. typus from the Chalk Group of the United Kingdom and the Hesseltal Formation of Germany.
Acrognathus is an extinct genus of ray-finned fish belonging to the order Aulopiformes. Although no extensive systematic analysis has been performed, it is tentatively placed with the greeneyes in the family Chlorophthalmidae, making it the oldest representative of that family.
Anomoeodus is an extinct genus of prehistoric marine ray-finned fish belonging to the family Pycnodontidae. This genus primarily lived during the mid-to-late Cretaceous period, ranging from the Albian to the very end of the Maastrichtian age, and possibly into the Danian. The first fossils of Anomoeodus were described by Louis Agassiz in 1833, although they were described under Pycnodus. Some studies have recovered it as a wastebasket taxon.
Ctenothrissa is a prehistoric genus of marine ray-finned fish in the order Ctenothrissiformes. It contains a number of species known from the Late Cretaceous of England and Lebanon.
Cylindracanthus is an extinct, enigmatic genus of marine ray-finned fish with fossils known throughout North America, Europe, Asia and Africa from the Late Cretaceous to the late Eocene, with potential Oligocene records and a possible Miocene record also known. It is exclusively known from its distinctive partial remains, which are long cylindrical bony spines that are usually considered rostrum fragments, as well as some associated teeth. These spines are abundant & widespread throughout this timespan, and are useful indicators of a nearshore marine environment, but the taxonomic identity of the fish is still highly uncertain and debated.
Enchodontidae is an extinct family of aulopiform ray-finned fish known from the mid-late Cretaceous. It contains two subfamilies with several genera, including the famous Enchodus, with great morphological disparity among members of the group.
The Dercetidae are an extinct family of aulopiform ray-finned fish that are known from the Late Cretaceous to the early Paleocene. They are among the many members of the diverse, extinct suborder Enchodontoidei, which were dominant during the Cretaceous.
Ichthyotringidae is an extinct family of aulopiform ray-finned fish known from the Early to Late Cretaceous. It is one of the Enchodontoidei, a diverse group of aulopiforms that were dominant marine fish during the Cretaceous.
Halecidae is an extinct family of aulopiform ray-finned fish known from the Cretaceous. It is one of the Enchodontoidei, a diverse group of aulopiforms that were dominant marine fish during the Cretaceous.
The Enchodontoidei are an extinct superorder of aulopiform fish known from the Early Cretaceous to the Eocene. They were among the dominant predatory marine fish groups in the Late Cretaceous, achieving a worldwide distribution. They were an extremely diverse group, with some developing fusiform body plans whereas others evolved elongated body plans with long beaks, superficially similar to eels and needlefish. They could also grow to very large sizes, as seen with Cimolichthys and Stratodus, the latter of which is the largest aulopiform known. Their most famous member is the widespread, abundant, and long-lasting genus Enchodus.