Cimolichthys Temporal range: | |
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C. nepaholica specimen, Denver Museum of Nature & Science | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Actinopterygii |
Order: | Aulopiformes |
Superfamily: | † Cimolichthyoidea |
Family: | † Cimolichthyidae Goody, 1969 |
Genus: | † Cimolichthys Leidy, 1857 |
Type species | |
†Cimolichthys levesiensis Leidy, 1857 | |
Species | |
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Synonyms | |
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Cimolichthys is an extinct genus of large predatory marine aulopiform fish known worldwide from the Late Cretaceous. It is the only member of the family Cimolichthyidae. [2] [3]
Cimolichthys was a large-sized nektonic aulopiform fish, making it related to modern lancetfish and lizardfish. Within the Aulopiformes, it is generally considered a member of the Enchodontoidei, a dominant group of predatory nektonic fish throughout much of the Cretaceous; [3] [4] however, some other treatments instead place it outside the Enchodontoidei and in a basal position as sister to the waryfishes, a small family of extant deep-sea aulopiformes. [5] Yet other studies have instead placed it as a sister to the extant daggertooths. [6]
The following species are known: [7]
C. anceps, C. gladiolus, C. semianceps, C. contracta, C. merillii, and C. sulcatus, all described by Cope (1872), are now considered synonymous with C. nepaholica. [3] [12] Indeterminate remains are known from the Maastrichtian of Niger & Brazil, and the Turonian of the Czech Republic. [7] [13]
Although the closest living relatives of Cimolichthys are lancetfish and lizardfish, the living animals would have resembled very large pikes. They could grow up to 1.5 to 2.0 metres (4.9 to 6.6 ft) meters long. Their bodies were covered by large, heavy scutes. Typical of this species are narrow lower jaws with several series of teeth. Remains of undigested fishes or squids have been found in collected specimens. Particularly well-preserved specimens are known from the Niobrara Formation of Kansas. [14] Their barbed teeth are distinctive, and can identify the presence of this genus even in formations where articulated remains are not otherwise known. [11]
Aulopiformes is a diverse order of marine ray-finned fish consisting of some 15 extant and several prehistoric families with about 45 genera and over 230 species. The common names grinners, lizardfishes and allies, or aulopiforms are sometimes used for this group. The scientific name means "Aulopus-shaped", from Aulopus + the standard fish order suffix "-formes". It ultimately derives from Ancient Greek aulós + Latin forma, the former in reference to the elongated shape of many aulopiforms.
Enchodus is an extinct genus of aulopiform ray-finned fish related to lancetfish and lizardfish. Species of Enchodus flourished during the Late Cretaceous, and there is some evidence that they may have survived to the Paleocene or Eocene; however, this may just represent reworked Cretaceous material.
Berycopsis is an extinct genus of marine ray-finned fish from the Late Cretaceous period. Fossils are known from England, Germany, and Lebanon. A potential specimen is known from the Czech Republic.
Belonostomus is a genus of prehistoric ray-finned fish that was described by Louis Agassiz in 1844. It is a member of the order Aspidorhynchiformes, a group of fish known for their distinctive elongated rostrums.
Apateodus is a genus of prehistoric marine ray-finned fish which was described by Woodward in 1901. It was a relative of modern lizardfish and lancetfish in the order Aulopiformes, and one of a number of prominent nektonic aulopiforms of Cretaceous marine ecosystems.
Apateopholis is an extinct genus of prehistoric marine ray-finned fish. It is the only member of the family Apateopholidae and contains a single species, A. laniatus, from the Cenomanian of Lebanon. It was a relative of modern lizardfish and lancetfish in the order Aulopiformes. At least one study has found it to be a potential sister genus to the enigmatic Yabrudichthys of the West Bank.
Argillichthys is an extinct marine lizardfish known from the Lower Eocene. It contains a single species, A. toombsi, from the Ypresian-aged London Clay Formation in England. It is known from a skull. It is thought to be a stem-member of Synodontidae.
Cyranichthys is an extinct marine aulopiform fish genus from Cenomanian-aged marine strata of what is now the Kipala formation in the Congo, Belgium and The Netherlands. The type species C. ornatissimus is known from Africa, while the newer discovered species, C. jagti, is known from Europe.
Caproberyx is an extinct genus of marine acanthomorph ray-finned fish, possibly a holocentrid, from the Late Cretaceous.
Cladocyclus is an extinct genus of marine ichthyodectiform fish from the middle Cretaceous. It was a predatory fish of about 1.20 metres (3.9 ft) in length.
Aulopopsis is an extinct genus of prehistoric marine ray-finned fish that lived during the lower Eocene. It is considered a relative of lizardfish in the order Aulopiformes, but its exact taxonomic placement is uncertain. Some authorities place it with the Aulopidae, while others place it with the Giganturoidae.
Aulolepis is an extinct genus of prehistoric marine ray-finned fish that lived during the upper Cenomanian and early Turonian. It contains a single species, A. typus from the Chalk Group of the United Kingdom and the Hesseltal Formation of Germany.
Acrognathus is an extinct genus of prehistoric bony fish belonging to the order Aulopiformes. Although no extensive systematic analysis has been performed, it is tentatively placed with the greeneyes in the family Chlorophthalmidae, making it the oldest representative of that family.
Anomoeodus is an extinct genus of prehistoric marine ray-finned fish belonging to the family Pycnodontidae. This genus primarily lived during the mid-to-late Cretaceous period, ranging from the Albian to the very end of the Maastrichtian age, and possibly into the Danian. The first fossils of Anomoeodus were described by Louis Agassiz in 1833, although they were described under Pycnodus. Some studies have recovered it as a wastebasket taxon.
Coelodus is an extinct genus of marine and possibly freshwater pycnodont fish. It contains only one definitive species, C. saturnusHeckel, 1854, from the Late Cretaceous of Slovenia. Other species from the Late Jurassic to the Eocene have also been attributed to this genus based on isolated dental elements, but their assignment to Coelodus is uncertain, and this genus likely represents a non-monophyletic wastebasket taxon. A potential diagnostic trait is a prearticular tooth row with three regular highly elongated teeth.
Enchodontidae is an extinct family of aulopiform fish known from the mid-late Cretaceous. It contains two subfamilies with several genera, including the famous Enchodus, with great morphological disparity among members of the group.
The Dercetidae are an extinct family of aulopiform fish that are known from the Late Cretaceous to the Paleocene. They are among the many members of the diverse, extinct suborder Enchodontoidei, which were dominant during the Cretaceous. Many genera evolved a very slender body plan with elongated jaws, closely converging on modern needlefish.
Ichthyotringidae is an extinct family of aulopiform fish known from the Early to Late Cretaceous. It is one of the Enchodontoidei, a diverse group of aulopiforms that were dominant marine fish during the Cretaceous.
Halecidae is an extinct family of aulopiform fish known from the Cretaceous. It is one of the Enchodontoidei, a diverse group of aulopiforms that were dominant marine fish during the Cretaceous.
The Enchodontoidei are an extinct superorder of aulopiform fish known from the Early Cretaceous to the Eocene. They were among the dominant predatory marine fish groups in the Late Cretaceous, achieving a worldwide distribution. They were an extremely diverse group, with some developing fusiform body plans whereas others evolved elongated body plans with long beaks, superficially similar to eels and needlefish. They could also grow to very large sizes, as seen with Cimolichthys and Stratodus, the latter of which is the largest aulopiform known. Their most famous member is the widespread, abundant, and long-lasting genus Enchodus.