| Clupavus Temporal range: | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| (unranked): | Otophysi |
| Family: | † Clupavidae |
| Genus: | † Clupavus Arambourg, 1950 |
| Type species | |
| †Clupavus maroccanus Arambourg, 1968 | |
| Species | |
Clupavus is an extinct genus of marine ray-finned fish that lived during the middle of the Cretaceous period. [3] It is known from North Africa, Europe, Brazil, and possibly North America.
It contains the following species:
Fossils of an indeterminate Clupavus species are abundant in the Albian-aged Pietraroja Plattenkalk of Italy, and the Cenomanian-aged Komen Limestone of Slovenia. [7] [8] A few specimens of a potential indeterminate species are also known from the Albian-aged Mowry Shale of Wyoming, USA. [9] [10]
The genus was initially described in 1950 with the species C. neocomiensis(Bassani, 1879) as its type species, based on a specimen from Morocco classified C. cf. neocomiensis. In 1968, the original C. neocomiensis was found to be synonymous with Leptolepis brodiei, so Clupavus was redefined with the newly-described C. maroccanus as the type species. [11] [12] The classification of C. brasiliensis in this genus has been questioned, as it differs from C. maroccanus in the morphology of the caudal skeleton. [13] [14] The species Casieroides yamangaensis and Chardonius longicaudatus from the Democratic Republic of the Congo were initially classified in this genus, but are now known to be unrelated. [4]
Initially described as a member of the Clupeiformes, [12] later studies have found the presence of a Weberian apparatus in members of this genus, indicating them to be basal otophysans. [4] [15] Relatives of Clupavus include Lusitanichthys and possibly Jhingrania . [16] [17]
Clupavus is one of the most common fossil fishes in the formations where it occurs in Europe. [5] [7] [8] They are especially common in the Hesseltal Formation of Germany, from around the Cenomanian-Turonian boundary. These German fossils indicate that Clupavus was an widespread, sardine-like shoaling fish, closely associated with cool waters from upwelling, that served as the base of the food chain in marine ecosystems of the Tethys Sea and pre-North Sea. Remains of Clupavus have been found as the fossilized stomach contents of medium-sized predatory fishes such as Bananogmius ornatus and possibly Protostomias maroccanus . [5]
Clupeiformes is the order of ray-finned fish that includes the herring family, Clupeidae, and the anchovy family, Engraulidae. The group includes many of the most important forage and food fish.
Ichthyodectiformes is an extinct order of marine stem-teleost ray-finned fish. The order is named after the genus Ichthyodectes, established by Edward Drinker Cope in 1870. Ichthyodectiforms are usually considered to be some of the closest relatives of the teleost crown group.
Cimolichthys is an extinct genus of large predatory marine aulopiform fish known worldwide from the Late Cretaceous. It is the only member of the family Cimolichthyidae.
Axelrodichthys is an extinct genus of mawsoniid coelacanth from the Cretaceous of Africa, North and South America, and Europe. Several species are known, the remains of which were discovered in the Lower Cretaceous (Aptian-Albian) of Brazil, North Africa, and possibly Mexico, as well as in the Upper Cretaceous of Morocco (Cenomanian), Madagascar and France. The Axelrodichthys of the Lower Cretaceous frequented both brackish and coastal marine waters while the most recent species lived exclusively in fresh waters. The French specimens are the last known fresh water coelacanths. Most of the species of this genus reached 1 metre to 2 metres in length. Axelrodichthys was named in 1986 by John G. Maisey in honor of the American ichthyologist Herbert R. Axelrod.
Apateodus is a genus of prehistoric marine ray-finned fish which was described by Woodward in 1901. It was a relative of modern lizardfish and lancetfish in the order Aulopiformes, and one of a number of prominent nektonic aulopiforms of Cretaceous marine ecosystems.
Bananogmius is an extinct genus of marine ray-finned fish that was found in what is now North America and Europe during the Late Cretaceous, from the Cenomanian to the Santonian. It lived in the Western Interior Seaway, which split North America in two during the Late Cretaceous, as well as the proto-North Sea of Europe.
Cladocyclus is an extinct genus of marine ichthyodectiform fish from the middle Cretaceous. It was a predatory fish of about 1.20 metres (3.9 ft) in length.
Chirocentrites is an extinct genus of marine ray-finned fish in the order Ichthyodectiformes. It contains a single species, C. coroninii, from the Late Cretaceous (Cenomanian) of Slovenia. A potential specimen is also known from the Albian-aged Pietraroja Plattenkalk of southern Italy, but it has been suggested that this specimen actually represents Cladocyclus.
Carsosaurus is a genus of extinct amphibious reptiles, in the mosasaur superfamily, containing only the species Carsosaurus marchesetti. It is known from a single individual that lived during the Upper Cretaceous in what is now Slovenia. The specimen is well-preserved, containing many different bones as well as some skin impressions and sternal cartilage. While more remains are needed to be certain, it is generally thought to belong to the Aigialosauridae. In life, it was an amphibious creature that spent most of its time on land, although its later relatives would become fully aquatic.
Aulolepis is an extinct genus of prehistoric marine ray-finned fish that lived during the upper Cenomanian and early Turonian. It contains a single species, A. typus from the Chalk Group of the United Kingdom and the Hesseltal Formation of Germany.
Araripichthys is an extinct genus of marine ray-finned fish that lived from the Aptian to Coniacian stages of the Cretaceous period. The genus is named after the Araripe Basin, where it was found in the Crato and Santana Formations. Other fossils of the genus have been found at Goulmima in Morocco, the Tlayua Formation of Mexico and the Apón Formation of Venezuela.
Anomoeodus is an extinct genus of prehistoric marine ray-finned fish belonging to the family Pycnodontidae. This genus primarily lived during the mid-to-late Cretaceous period, ranging from the Albian to the very end of the Maastrichtian age, and possibly into the Danian. The first fossils of Anomoeodus were described by Louis Agassiz in 1833, although they were described under Pycnodus. Some studies have recovered it as a wastebasket taxon.
Coelodus is an extinct genus of marine and possibly freshwater pycnodont fish. It contains only one definitive species, C. saturnusHeckel, 1854, from the Late Cretaceous of Slovenia. Other species from the Late Jurassic to the Eocene have also been attributed to this genus based on isolated dental elements, but their assignment to Coelodus is uncertain, and this genus likely represents a non-monophyletic wastebasket taxon. A potential diagnostic trait is a prearticular tooth row with three regular highly elongated teeth.
Caeus is an extinct genus of prehistoric marine ray-finned fish, closely related to the modern milkfish. It contains a single species, C. leopoldi from the Early Cretaceous of the Pietraroja Plattenkalk, Italy. It is one of the largest teleosts known from the Pietraroja formation, and is known by only a single specimen.
Pycnodontidae is an extinct family of ray-finned fishes, ranging from the Jurassic period until the Eocene. It was the largest and most derived family of the successful Mesozoic fish order Pycnodontiformes, and the only member of it to survive into the Cenozoic.
Armigatus is an extinct genus of marine clupeomorph fishes belonging to the order Ellimmichthyiformes. These fishes lived in the Cretaceous ; their fossil remains have been found in Mexico, Europe, the Middle East and North Africa, suggesting the genus ranged across the Tethys Sea.
This list of fossil fishes described in 2020 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes of every kind that were described during the year 2020, as well as other significant discoveries and events related to paleoichthyology that occurred in 2020.
The Hesseltal Formation or Blackcoloured Formation is a Late Cretaceous geological formation from northern Germany. It consists of lithified marls and limestone, with a unique series of black shales deposited in anoxic conditions during the Cenomanian-Turonian boundary event.
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