Cordylidae | |
---|---|
| |
Tropical girdled lizard, Cordylus tropidosternum | |
Scientific classification ![]() | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Reptilia |
Order: | Squamata |
Superfamily: | Cordyloidea |
Family: | Cordylidae Fitzinger, 1826 |
Diversity | |
10 genera (see text) |
Cordylidae is a family of small- to medium-sized lizards that occur in southern and eastern Africa. They are commonly known as girdled lizards, spinytail lizards, or girdle-tail lizards. [1] [2]
Cordylidae is closely related to the family Gerrhosauridae, occurring in Africa and Madagascar. These two scientific families of lizards, known as Cordyliformes or Cordyloidea, are sometimes combined into a larger concept of Cordylidae. Recent molecular analyses confirm the clade made up of Cordylidae and Gerrhosauridae (Cordyloidea) and place it in a larger clade including Xantusiidae (Cordylomorpha Vidal & Hedges, 2009). [3]
Girdled lizards are diurnal and insectivorous. They are terrestrial, mostly inhabiting crevices in rocky terrain, although at least one species digs burrows and another lives under exfoliating bark on trees. They have flattened heads and bodies, and are distinguished by a heavy armour of osteoderms and large, rectangular, scales, arranged in regular rows around the body and tail. Many species have rings of spines on the tail, that aid in wedging the animal into sheltering crevices, and also in dissuading predators. [4]
Most species have four limbs, but those in the genus Chamaesaura are almost entirely limbless, with only tiny spikes in place of the hind limbs. The family includes both egg-laying and ovoviviparous species. [4]
List of genera | |||||
Genus | Image | Type species | Taxon author | Common name | Species |
---|---|---|---|---|---|
Chamaesaura | ![]() C. anguina | C. anguina (Linnaeus, 1758) | Schneider, 1801 | Grass lizards | 5 |
Cordylus | ![]() C. niger | C. cordylus (Linnaeus, 1758) | Laurenti, 1768 | Girdled lizards | 21 |
Hemicordylus | ![]() H. capensis | H. capensis (Smith, 1838) | Smith, 1838 | Cliff lizards | 2 |
Karusasaurus | ![]() K. polyzonus | K. polyzonus (Smith, 1838) | Stanley, Bauer, Jackman, Branch & Mouton, 2011 | Karusa lizards | 2 |
Namazonurus | ![]() N. peersi | N. pustulatus (W. Peters, 1862) | Stanley, Bauer, Jackman, Branch & Mouton, 2011 | Namaqua girdled lizards | 5 |
Ninurta | ![]() N. coeruleopunctatus | N. coeruleopunctatus (Methuen & Hewitt, 1913) | Stanley, Bauer, Jackman, Branch & Mouton, 2011 | Blue-spotted girdled lizard | 1 |
Ouroborus | ![]() O. cataphractus | O. cataphractus (F. Boie, 1828) | Stanley, Bauer, Jackman, Branch & Mouton, 2011 | Armadillo girdled lizard | 1 |
Platysaurus | P. relictus | P. capensis Smith, 1844 | Smith, 1844 | Flat lizards | 16 |
Pseudocordylus | ![]() P. subviridis | P. microlepidotus (Cuvier, 1829) | Smith, 1838 | Crag lizards | 6 |
Smaug | S. giganteus | S. giganteus (Smith, 1844) | Stanley, Bauer, Jackman, Branch & Mouton, 2011 | Girdled lizards | 9 |
The Lepidosauria is a subclass or superorder of reptiles, containing the orders Squamata and Rhynchocephalia. Squamata includes snakes, lizards, and amphisbaenians. Squamata contains over 9,000 species, making it by far the most species-rich and diverse order of reptiles in the present day. Rhynchocephalia was a formerly widespread and diverse group of reptiles in the Mesozoic Era. However, it is represented by only one living species: the tuatara, a superficially lizard-like reptile native to New Zealand.
Amphisbaenia is a group of usually legless squamates, comprising over 200 extant species. Amphisbaenians are characterized by their long bodies, the reduction or loss of the limbs, and rudimentary eyes. As many species have a pink body and scales arranged in rings, they have a superficial resemblance to earthworms. While the genus Bipes retains forelimbs, all other genera are limbless. Although superficially similar to the snakes and blind lizards, recent phylogenetic studies suggest that they are most closely related to wall lizards of the family Lacertidae. Amphisbaenians are widely distributed, occurring in North America, Europe, Africa, South America, Western Asia and the Caribbean. They are not found east of the Caspian Sea. Most species are less than 6 inches (150 mm) long.
The Gerrhosauridae are a family of lizards native to Africa and Madagascar.
The genus Cordylus includes a wide variety of species of small to medium spiny lizards from Africa, collectively called girdle-tailed lizards or girdled lizards. All are diurnal and ovoviviparous. Most species are rupicolous (rock-dwelling), while a few species are arboreal or live in burrows. They defend themselves with osteoderms and by quickly retreating into rock crevices or burrows. Many species live in groups, and males defend territories.
Ophidia is a group of squamate reptiles including modern snakes and reptiles more closely related to snakes than to other living groups of lizards.
The armadillo girdled lizard, also commonly known as the armadillo lizard, the armadillo spiny-tailed lizard, and the golden-armadillo lizard, is a species of lizard in the family Cordylidae. The species is endemic to desert areas along the western coast of South Africa. In 2011, it was moved to its own genus based on molecular phylogeny, but formerly it was included in the genus Cordylus.
Autarchoglossa is a clade of squamates that includes skinks, anguimorphs, snakes, and relatives. Autarchoglossa is supported as a monophyletic grouping by morphological features in living and extinct lizards and snakes. Some phylogenetic analyses based on molecular features such as DNA sequences in living squamates do not support Autarchoglossa.
Legless lizard may refer to any of several groups of lizards that have independently lost limbs or reduced them to the point of being of no use in locomotion. It is the common name for the family Pygopodidae,. These lizards are often distinguishable from snakes on the basis of one or more of the following characteristics: possessing eyelids, possessing external ear openings, lack of broad belly scales, notched rather than forked tongue, having two more-or-less-equal lungs, and/or having a very long tail.
The sungazer, also known as the giant girdled lizard, giant dragon lizard, or giant zonure, is the largest species of the Cordylidae, a family of lizards from sub-Saharan Africa. This threatened species is endemic to Highveld grasslands in the interior of South Africa. In 2011, it was assigned to the new genus Smaug, along with seven other species previously belonging to the genus Cordylus, based on a comprehensive molecular phylogeny of the Cordylidae.
Scincomorpha is an infraorder and clade of lizards including skinks (Scincidae) and their close relatives. These include the living families Cordylidae, Gerrhosauridae, and Xantusiidae, as well as many extinct taxa. Other roughly equivalent terms include the suborder Scinciformata, or the superfamily Scincoidea, though different authors use these terms in a broader or more restricted usage relative to true skinks. They first appear in the fossil record about 170 million years ago, during the Jurassic period. The phylogeny below follows that of Alifanov in 2016.
The Limpopo girdled lizard, also known commonly as Jones's armadillo lizard and Jones's girdled lizard, is a species of lizard in the family Cordylidae. The species is endemic to Southern Africa.
An isolated population of the Rhodesian girdled lizard from granite outcrops in montane grassland of northern Malawi was recently redescribed as Cordylus nyikae. Unlike the Rhodesian girdled lizard, the head shields of this species are very rugose, the nostrils are pierced in the lower posterior corner of the nasal scales, and the dorsals do not have a serrated posterior margin. The dorsal coloration is dark brown to gray-brown with lighter spots. The tail and upper lips are light brown. The belly is buff. Its common name is Nyika girdled lizard.
The Rhodesian girdled lizard is one of three species of very flattened girdled lizards from Zimbabwe, Malawi, and Mozambique. The other two species are Cordylus nyikae and Cordylus meculae. They are found living under stones in rock outcrops where they feed on small arthropods. Adults are 60–90 mm long from snout to vent. The tail length is approximately half the total length.
Warren's girdled lizard is a species of relatively large, flattened lizard in the family Cordylidae. The species is native to Southern Africa.
The Anguimorpha is a suborder of squamates. The group was named by Fürbringer in 1900 to include all autarchoglossans closer to Varanus and Anguis than Scincus. These lizards, along with iguanians and snakes, constitute the proposed "venom clade" Toxicofera of all venomous reptiles. The oldest definitive record of the group is Dorsetisaurus from the Late Jurassic of Europe and North America.
The Lacertoidea is a group of squamate reptiles that includes the Lacertidae, Teiidae, Gymnophthalmidae, and Amphisbaenia. The finding from molecular phylogenetic studies that the burrowing Amphisbaenia were nested in a clade with the lizard forms led Vidal & Hedges (2005) to propose a new name for the group based on shared morphogical characters, Laterata, "referring to the presence of tile-like scales that form the rings in Amphisbaenia, and are also present ventrally in Lacertiformata and Teiformata".
Namazonurus campbelli, commonly known as Campbell's girdled lizard, is a species of lizard in the family Cordylidae. A small girdled lizard, N. campbelli is endemic to Namibia. It is often mistaken for the more common Herero girdled lizard, N. pustulatus, as they both are similar in size and have flattened bodies. N. campbelli lives in rock crevices and cracks on dry mountain slopes
Smaug is a genus of lizards in the family Cordylidae. The genus Smaug is a group of species of spiny southern African lizards, separated from the genus Cordylus in 2011 on the basis of a comprehensive molecular phylogeny of the Cordylidae. The type species is the giant girdled lizard, S. giganteus.
Van Dam's girdled lizard is a species of lizard in the family Cordylidae. The species is endemic to South Africa.
Smaug breyeri, also known commonly as the Waterberg dragon lizard or the Waterberg girdled lizard, is a species of lizard in the family Cordylidae. The species is endemic to South Africa.