Cyrilavis Temporal range: Early Eocene, | |
---|---|
Skull of C. colburnorum | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Family: | † Halcyornithidae |
Genus: | † Cyrilavis Martin, 2010 |
Type species | |
†Primobucco olsoni (Feduccia & Martin, 1976) |
Cyrilavis is an extinct genus of halcyornithid bird from the Early Eocene Fossil Butte Member of the Green River Formation, United States. The genus contains two species, Cyrilavis olsoni and Cyrilavis colburnorum. [1]
A complete skeleton of a bird from the Green River Formation was described as "Primobucco" olsoni in 1976 from the speciment GSATC 217 as a relative to the Bucconidae based on its zygodactyl foot configuration. The specific ephitet olsoni referred to Storrs L. Olson. [1] [2] In 2002, the specimen was recognised as having psittacine affinities by Mayr. [3] In 2010 the genus Cyrilavis was erected to accomdoate the halcyornithid, since it was no longer thought to be closely related to the Bucconidae, with the new name Cyrilavis olsoni. The generic name Cyrilavis is in honour of Cyril Walker, a notable paleontologist of early Cenozoic birds. [4] In 2011, Ksepka et al. described a new species, Cyrilavis colburnorum, with the holotype FMNH PA 754, a nearly complete skeleton. Another specimen of C. colburnorum, FMNH PA 766, preserved both feet and much of the legs and wings. A third specimen is not referred to either species. [1]
Cyrilavis was a relatively large-headed bird. The beak is about half of the total length of the head, and is dorsoventrally expanded. The lower half of the beak remains shallow while the top half is comparatively tall. As in all other halcyornithids and in the psittacopasseran Psittacopes , the manus is much shorter than the ulna. This is the opposite of the condition found in extant parrots. [1]
Cyrilavis olsoni, whose holotype preserves feathering, shows a halo of feathers around the head, interpreted as a crest. [1]
The holotype of Cyrilavis was initially referred to the Primobucconidae, a family of stem puffbirds. Later revisions resulted in the genus' interpretation as a stem parrot. Recent analyses have found that the Halcyornithidae, to which Cyrilavis belongs, diverged before the split between parrots and passerines, meaning it is equally closely related to both orders. [5]
Palaeotis is a genus of paleognath birds from the middle Eocene epoch of central Europe. One species is known, Palaeotis weigelti. The holotype specimen is a fossil tarsometatarsus and phalanx. Lambrect (1928) described it as an extinct bustard, and gave it its consequent name. After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that Palaeotis is a palaeognath and assigned it to the same order as ostriches; the Struthioniformes. In 2021, it was considered a member of the family Paleotididae alongside Galligeranoides from the Early Eocene of France, which were found to be basal members of the Struthioniformes.
"Precursor" is a controversial prehistoric bird genus from the Early Eocene. It was established based on fossils found in England, including in the famous London Clay deposits. Three species are included in the genus: "P." parvus, the type species, "P." magnus, and "P." litorum, all named by Colin Harrison and Cyril Walker in 1977.
Rhynchaeites is an extinct genus of wading bird, a stem-group threshkiornithid, from the Eocene. It is one of the oldest members of the ibis family known from fossil remains.
Mopsitta tanta is an extinct bird of uncertain taxonomic position from the Early Eocene of Denmark; its remains were recovered from the Fur Formation. So far, the holotype and only known specimen is a single humerus bone of rather large size. Although the phylogenetic position of genus is unclear, it was initially presumed to be phylogenetically closer to Recent Psittacidae than to other known Palaeogene psittaciforms and may, therefore, represent the oldest known crown-group parrot.
Eurofluvioviridavis is a genus of extinct birds from the Middle Eocene Messel Pit, Germany. It contains a single species, Eurofluvioviridavis robustipes. It is related to Avolatavis and Vastanavis, other members of the family Vastanavidae.
Amitabha urbsinterdictensis is an ancient bird from the Middle Eocene in North America. One specimen has been found to date. Bonnie Gulas-Wroblewski and Anton Wroblewski described and named it in 2002.
Avolatavis is an extinct genus of stem-parrot (pan-psittaciform) or a member of the stem group of Psittacopasseres, known from the early Eocene Fossil Butte Member of the Green River Formation of Wyoming, United States, and from the London Clay of Walton-on-the-Naze. It was first named by Daniel T. Ksepka and Julia A. Clarke in 2012 and the type species is Avolatavis tenens. Gerald Mayr and Andrew C. Kitchener described the second species, A. europaeus, in 2023. Mayr and Kitchener assigned Avolatavis to the family Vastanavidae, which might be early diverging stem group presentatives of Pan-Psittaciformes or stem group representatives of Psittacopasseres.
Messelasturidae is an extinct family of birds known from the Eocene of North America and Europe. Their morphology is a mosaic that in some aspects are very similar to modern hawks and falcons, but in others are more similar to parrots. Initially interpreted as stem-owls, more recent studies have suggested a closer relationship to parrots and passerines. Their ecology is enigmatic.
Vastanavidae is an extinct family of birds related to parrots and passerine birds. They are known from fossils from Eocene sites in India, Europe, and North America. The vastanavids resemble parrots and the extinct parrot relative Quercypsitta in their morphology, including the partially zygodactyl foot, in which two toes could face opposite the other two.
Vegaviidae is an extinct family of birds of uncertain phylogenetic placement, which existed during the Late Cretaceous and possibly the Paleocene. Definitive fossils attributed to the family have only been found in Antarctica, though other fossils from the Southern Hemisphere including Chile and New Zealand may represent this group. The putative Campanian vegaviid from Canada known as Maaqwi, more likely belongs to the Procellariformes.
Halcyornithidae is an extinct family of telluravian birds thought to be related to the Psittaciformes (parrots), Passeriformes (songbirds), and to the extinct Messelasturidae. Halcyornithids have been found in various Eocene formations in Europe and North America. Widespread and diverse in the Early Eocene of North America and Europe, halcyornithids are not found in locales later than the Middle Eocene. Halcyornithids were small, arboreal birds with zygodactyl feet, with two toes facing forwards and two facing back, a trait shared with other tree-dwelling families of Eocene birds like the Zygodactylidae and the messelasturids. The skull of halcyornithids features a ridge of bone above the eye called the supraorbital process, similar to birds of prey. The relationships of the halcyornithids to other birds remain uncertain. Halcyornithids have been proposed as relatives to owls and as a lineage closer to parrots than to songbirds. Most recently, halcyornithids have been identified as the sister group of the clade including parrots and songbirds. It is also possible that Halcyornithidae is paraphyletic with respect to the Messelasturidae.
Morsoravis is an extinct genus of neoavian bird from the Early Eocene Fur Formation of Denmark. It contains a single named species, Morsoravis sedilis. Fossils of Morsoravis have also been found in the Green River Formation of Wyoming and possibly the Nanjemoy Formation of Virginia.
Danielsraptor is an extinct genus of masillaraptorid bird from the Early Eocene (Ypresian) Walton Member of the London Clay Formation in Essex, United Kingdom. The genus contains a single species, D. phorusrhacoides, known from a partial skeleton.
Halcyornis is an extinct genus of halcyornithid bird. It is known from a single neurocranium from the Early Eocene London Clay of southeastern England, and contains a single species, Halcyornis toliapicus. The type specimen of Halcyornis was the first fossil bird to be scientifically named.
Pulchrapollia is an extinct genus of halcyornithid bird from the Early Eocene London Clay of Walton-on-the-Naze, United Kingdom and the Nanjemoy Formation of Virginia, United States. The genus contains three species, Pulchrapollia gracilis, Pulchrapollia tenuipes and Pulchrapollia eximia.
Pseudasturides is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany, and possibly the Isle of Sheppey, United Kingdom. The genus is represented by a single species, Pseudasturides macrocephalus.
Serudaptus is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany. The genus contains one species, Serudaptus pohli, and is known for long, raptorial claws on its zygodactyl feet.
Eofringillirostrum is an extinct species of bird known from the Early Eocene Green River Formation of the Western United States and from the Messel Pit in Germany. The genus contains two species, Eofringillirostrum boudreauxi and Eofringillirostrum parvulum. They are the earliest known fossil birds to have a finch-like beak.
Piscator tenuirostris is an extinct species of cormorant-like bird, the only known species in the genus Piscator.
Paakniwatavis is a genus of extinct waterfowl from the early Eocene Green River Formation of Wyoming, United States. The genus contains a single species, P. grandei, known from a partial skeleton with a skull.