Cyrilavis

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Cyrilavis
Temporal range: Early Eocene, 51.6  Ma
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Parrot Green River.jpg
Skull of C. colburnorum
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Family: Halcyornithidae
Genus: Cyrilavis
Martin, 2010
Type species
Primobucco olsoni
(Feduccia & Martin, 1976)

Cyrilavis is an extinct genus of halcyornithid bird from the Early Eocene Fossil Butte Member of the Green River Formation, United States. The genus contains two species, Cyrilavis olsoni and Cyrilavis colburnorum. [1]

Contents

Discovery and naming

A complete skeleton of a bird from the Green River Formation was described as "Primobucco" olsoni in 1976 from the speciment GSATC 217 as a relative to the Bucconidae based on its zygodactyl foot configuration. The specific ephitet olsoni referred to Storrs L. Olson. [1] [2] In 2002, the specimen was recognised as having psittacine affinities by Mayr. [3] In 2010 the genus Cyrilavis was erected to accomdoate the halcyornithid, since it was no longer thought to be closely related to the Bucconidae, with the new name Cyrilavis olsoni. The generic name Cyrilavis is in honour of Cyril Walker, a notable paleontologist of early Cenozoic birds. [4] In 2011, Ksepka et al. described a new species, Cyrilavis colburnorum, with the holotype FMNH PA 754, a nearly complete skeleton. Another specimen of C. colburnorum, FMNH PA 766, preserved both feet and much of the legs and wings. A third specimen is not referred to either species. [1]

Description

Cyrilavis was a relatively large-headed bird. The beak is about half of the total length of the head, and is dorsoventrally expanded. The lower half of the beak remains shallow while the top half is comparatively tall. As in all other halcyornithids and in the psittacopasseran Psittacopes , the manus is much shorter than the ulna. This is the opposite of the condition found in extant parrots. [1]

Cyrilavis olsoni, whose holotype preserves feathering, shows a halo of feathers around the head, interpreted as a crest. [1]

Classification

The holotype of Cyrilavis was initially referred to the Primobucconidae, a family of stem puffbirds. Later revisions resulted in the genus' interpretation as a stem parrot. Recent analyses have found that the Halcyornithidae, to which Cyrilavis belongs, diverged before the split between parrots and passerines, meaning it is equally closely related to both orders. [5]

Related Research Articles

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The puffbirds and their relatives in the near passerine family Bucconidae are tropical tree-dwelling insectivorous birds that are found from South America up to Mexico. Together with their closest relatives, the jacamars, they form a divergent lineage within the order Piciformes, though the two families are sometimes elevated to a separate order Galbuliformes. Lacking the iridescent colours of the jacamars, puffbirds are mainly brown, rufous or grey, with large heads, large eyes, and flattened bills with a hooked tip. Their loose, abundant plumage and short tails makes them look stout and puffy, giving rise to the English name of the family. The species range in size from the rufous-capped nunlet, at 13 cm (5.1 in) and 14 g (0.49 oz), to the white-necked puffbird, at up to 29 cm (11 in) and 106 g (3.7 oz).

<i>Palaeotis</i> Extinct genus of birds

Palaeotis is a genus of paleognath birds from the middle Eocene epoch of central Europe. One species is known, Palaeotis weigelti. The holotype specimen is a fossil tarsometatarsus and phalanx. Lambrect (1928) described it as an extinct bustard, and gave it its consequent name. After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that Palaeotis is a palaeognath and assigned it to the same order as ostriches; the Struthioniformes. In 2021, it was considered a member of the family Paleotididae alongside Galligeranoides from the Early Eocene of France, which were found to be basal members of the Struthioniformes.

"Precursor" is a controversial prehistoric bird genus from the Early Eocene. It was established based on fossils found in England, including in the famous London Clay deposits. Three species are included in the genus: "P." parvus, the type species, "P." magnus, and "P." litorum, all named by Colin Harrison and Cyril Walker in 1977.

Mopsitta tanta is an extinct bird of uncertain taxonomic position from the Early Eocene of Denmark; its remains were recovered from the Fur Formation. So far, the holotype and only known specimen is a single humerus bone of rather large size. Although the phylogenetic position of genus is unclear, it was initially presumed to be phylogenetically closer to Recent Psittacidae than to other known Palaeogene psittaciforms and may, therefore, represent the oldest known crown-group parrot.

Psittacopes is an extinct genus of bird from Middle Eocene. One species is recorded from Messel, Germany, and other three possible species are from London Clay, England, one named ?Psittacopes occidentalis in 2022, and the other two unnamed. Its phylogenetic placement within Aves is uncertain; it was originally interpreted as a parrot, but the phylogenetic analysis conducted by Mayr (2015) recovered it as more closely related to the passerines and the extinct family Zygodactylidae.

Eurofluvioviridavis is a genus of extinct primitive birds from the Middle Eocene Messel Pit, Germany. It contains a single species, Eurofluvioviridavis robustipes. It is related to Avolatavis and Vastanavis, other members of the family Vastanavidae.

Amitabha urbsinterdictensis is an ancient bird from the Middle Eocene in North America. One specimen has been found to date. Bonnie Gulas-Wroblewski and Anton Wroblewski described and named it in 2002.

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<span class="mw-page-title-main">Messelasturidae</span> Extinct family of birds

Messelasturidae is an extinct family of birds known from the Eocene of North America and Europe. Their morphology is a mosaic that in some aspects are very similar to modern hawks and falcons, but in others are more similar to parrots. Initially interpreted as stem-owls, more recent studies have suggested a closer relationship to parrots and passerines. Their ecology is enigmatic.

Vastanavidae is an extinct family of birds related to parrots and passerine birds. They are known from fossils from Eocene sites in India, Europe, and North America. The vastanavids resemble parrots and the extinct parrot relative Quercypsitta in their morphology, including the partially zygodactyl foot, in which two toes could face opposite the other two.

<span class="mw-page-title-main">Halcyornithidae</span> Extinct family of birds

Halcyornithidae is an extinct family of telluravian birds thought to be related to the Psittaciformes (parrots), Passeriformes (songbirds), and to the extinct Messelasturidae. Halcyornithids have been found in various Eocene formations in Europe and North America. Widespread and diverse in the Early Eocene of North America and Europe, halcyornithids are not found in locales later than the Middle Eocene. Halcyornithids were small, arboreal birds with zygodactyl feet, with two toes facing forwards and two facing back, a trait shared with other tree-dwelling families of Eocene birds like the Zygodactylidae and the messelasturids. The skull of halcyornithids features a ridge of bone above the eye called the supraorbital process, similar to birds of prey. The relationships of the halcyornithids to other birds remain uncertain. Halcyornithids have been proposed as relatives to owls and as a lineage closer to parrots than to songbirds. Most recently, halcyornithids have been identified as the sister group of the clade including parrots and songbirds. It is also possible that Halcyornithidae is paraphyletic with respect to the Messelasturidae.

Morsoravis is an extinct genus of neoavian bird from the Early Eocene Fur Formation of Denmark. It contains a single named species, Morsoravis sedilis. Fossils of Morsoravis have also been found in the Green River Formation of Wyoming and possibly the Nanjemoy Formation of Virginia.

Danielsraptor is an extinct genus of masillaraptorid bird from the Early Eocene (Ypresian) Walton Member of the London Clay Formation in Essex, United Kingdom. The genus contains a single species, D. phorusrhacoides, known from a partial skeleton.

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Pulchrapollia is an extinct genus of halcyornithid bird from the Early Eocene London Clay of Walton-on-the-Naze, United Kingdom and the Nanjemoy Formation of Virginia, United States. The genus contains three species, Pulchrapollia gracilis, Pulchrapollia tenuipes and Pulchrapollia eximia.

Pseudasturides is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany, and possibly the Isle of Sheppey, United Kingdom. The genus is represented by a single species, Pseudasturides macrocephalus.

<i>Serudaptus</i> Extinct genus of birds

Serudaptus is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany. The genus contains one species, Serudaptus pohli, and is known for long, raptorial claws on its zygodactyl feet.

Eofringillirostrum is an extinct species of bird known from the Early Eocene Green River Formation of the Western United States and from the Messel Pit in Germany. The genus contains two species, Eofringillirostrum boudreauxi and Eofringillirostrum parvulum. They are the earliest known fossil birds to have a finch-like beak.

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References

  1. 1 2 3 4 5 Ksepka, Daniel T.; Clarke, Julia A.; Grande, Lance (2011). "Stem Parrots (Aves, Halcyornithidae) from the Green River Formation and a Combined Phylogeny of Pan-Psittaciformes". Journal of Paleontology. 85 (5): 835–852. doi:10.1666/10-108.1. ISSN   0022-3360.
  2. Feduccia A, Martin LD. 1976. The Eocene zygodactyl birds of North America (Aves: Piciformes). Smithsonian Contributions to Paleobiology 27: 101–110
  3. MAYR, GERALD (2002). "On the osteology and phylogenetic affinities of the Pseudasturidae–Lower Eocene stem-group representatives of parrots (Aves, Psittaciformes)". Zoological Journal of the Linnean Society. 136 (4): 715–729. doi: 10.1046/j.1096-3642.2002.00042.x . ISSN   1096-3642.
  4. Larry D. MARTIN . PALEOGENE AVIFAUNA OF THE HOLARCTIC [J]. Vertebrata Palasiatica, 2010, 48(4): 367-374.
  5. Mayr, Gerald (2022). Paleogene fossil birds. Fascinating life sciences (2nd ed.). Cham: Springer. ISBN   978-3-030-87644-9.