Eurofluvioviridavis Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Family: | † Vastanavidae |
Genus: | † Eurofluvioviridavis Mayr, 2005 |
Type species | |
†Eurofluvioviridavis robustipes (Mayr, 2005) | |
Synonyms | |
Eurofluvioviridavis is a genus of extinct birds from the Middle Eocene Messel Pit, Germany. It contains a single species, Eurofluvioviridavis robustipes. It is related to Avolatavis and Vastanavis , other members of the family Vastanavidae. [1] [2]
The holotype of Eurofluvioviridavis, SMNK.PAL.3835, was found in the lower middle Eocene Messel Pit, near Darmstadt, Germany, in the MP 11 Mammal Paleogene zone. The fossil consists of a well-preserved skeleton. An additional counter-slab of the holotype exists, but was not studied in the genus' description. [1]
Eurofluvioviridavis was scientifically described in Mayr (2005) as a bird similar to Fluvioviridavis from the Green River Formation in Wyoming. The generic name Eurofluvioviridavis refers to a similarity to the aforementioned taxon as well as the fossil's European provenance. The specific ephitet, robustipes, is from Latin robustus, meaning robust, and pes, meaning foot, in reference to strongly built feet. [1]
Eurofluvioviridavis had a wide, flattened beak, resembling that of tyrant flycatchers. It is as long as the rest of the head, and is constant in width in the rear half, tapering to a point in the front half. The nostrils are large. The palate is not heavily ossified, unlike in some extant birds with wide, flat beaks, and there are no basipterygoid processes. There are eight caudal vertebrae and the pygostyle is narrow. The coracoid possesses a foramen for the supracoracoid nerve. The lateral process of the coracoid is a small, hooklike structure. The humerus is proportionally falcon-like, with a large deltapectoral crest and a curved shaft. The ulna is longer than the humerus, with eight to thirteen quill knobs indicating a relatively low number of secondary feathers. The carpometacarpus is relatively long, and the phalanx of the alula has a poorly developed claw. The femur is short and stout, while the fibula is moderate in proportions. [1]
The tarsometatarsus and feet of Eurofluvioviridavis resemble those of Avolatavis and Vastanavis. The tarsometatarsus is stout. The feet show a semi-zygodactyl arrangement, and the ungual phalanges are raptor-like, possibly indicating that Eurofluvioviridavis could grasp with the foot. [3]
Cladistic analyses support a relationship between the genera Vastanavis, Avolatavis, and Eurofluvioviridavis in the family Vastanavidae. Their position relative to modern birds is not clear; it is possible that the vastanavids are stem-group relatives of the parrots, or that they are part of the stem group of the Psittacopasseres. These hypotheses are supported by morphological analysis and morphological analysis with molecular constraints, respectively. Eurofluvioviridavis, like other vastanavids and other psittacopasserean outgroups like the Halcyornithidae and the Messelasturidae, possess short tarsometatarsi. This is in the contrast to the long tarsometatarsi plesiomorphic to the Australaves, the clade including falcons, parrots, and songbirds, which appears to indicate that a short, stocky tarsometatarsus was plesiomorphic to Pan-Psittacopasseres, the total group including parrots, songbirds, and their stem-lineage representatives. [4]
Avolatavis has not been differentiated from Eurofluvioviridavis, meaning it is possible that the former taxon is a junior synonym. However, a non-overlap of important fossil material means that it is not currently possible to assess the status of these genera. [4]
Palaeotis is a genus of paleognath birds from the middle Eocene epoch of central Europe. One species is known, Palaeotis weigelti. The holotype specimen is a fossil tarsometatarsus and phalanx. Lambrect (1928) described it as an extinct bustard, and gave it its consequent name. After a suggestion by Storrs L. Olson, a review of the type specimen and the referral of several other fossils by Houde and Haubold (1987) concluded that Palaeotis is a palaeognath and assigned it to the same order as ostriches; the Struthioniformes. In 2021, it was considered a member of the family Paleotididae alongside Galligeranoides from the Early Eocene of France, which were found to be basal members of the Struthioniformes.
Rhynchaeites is an extinct genus of wading bird, a stem-group threshkiornithid, from the Eocene. It is one of the oldest members of the ibis family known from fossil remains.
Avolatavis is an extinct genus of stem-parrot (pan-psittaciform) or a member of the stem group of Psittacopasseres, known from the early Eocene Fossil Butte Member of the Green River Formation of Wyoming, United States, and from the London Clay of Walton-on-the-Naze. It was first named by Daniel T. Ksepka and Julia A. Clarke in 2012 and the type species is Avolatavis tenens. Gerald Mayr and Andrew C. Kitchener described the second species, A. europaeus, in 2023. Mayr and Kitchener assigned Avolatavis to the family Vastanavidae, which might be early diverging stem group presentatives of Pan-Psittaciformes or stem group representatives of Psittacopasseres.
Messelasturidae is an extinct family of birds known from the Eocene of North America and Europe. Their morphology is a mosaic that in some aspects are very similar to modern hawks and falcons, but in others are more similar to parrots. Initially interpreted as stem-owls, more recent studies have suggested a closer relationship to parrots and passerines. Their ecology is enigmatic.
Vastanavidae is an extinct family of birds related to parrots and passerine birds. They are known from fossils from Eocene sites in India, Europe, and North America. The vastanavids resemble parrots and the extinct parrot relative Quercypsitta in their morphology, including the partially zygodactyl foot, in which two toes could face opposite the other two.
Halcyornithidae is an extinct family of telluravian birds thought to be related to the Psittaciformes (parrots), Passeriformes (songbirds), and to the extinct Messelasturidae. Halcyornithids have been found in various Eocene formations in Europe and North America. Widespread and diverse in the Early Eocene of North America and Europe, halcyornithids are not found in locales later than the Middle Eocene. Halcyornithids were small, arboreal birds with zygodactyl feet, with two toes facing forwards and two facing back, a trait shared with other tree-dwelling families of Eocene birds like the Zygodactylidae and the messelasturids. The skull of halcyornithids features a ridge of bone above the eye called the supraorbital process, similar to birds of prey. The relationships of the halcyornithids to other birds remain uncertain. Halcyornithids have been proposed as relatives to owls and as a lineage closer to parrots than to songbirds. Most recently, halcyornithids have been identified as the sister group of the clade including parrots and songbirds. It is also possible that Halcyornithidae is paraphyletic with respect to the Messelasturidae.
Oligocolius is an unusual genus of extinct mousebird from the early to late Oligocene epoch of Germany. Oligocolius is known from two species, the type species O. brevitarsus and O. psittacocephalon. Oligocolius is a member of the family of modern mousebirds (Coliidae) and is broadly similar to them in shape. However, unlike modern mouse birds the skull and beak of Oligocolius closely resembles those of parrots, and appears to be specialised for a distinct lifestyle not found in living mousebirds.
Morsoravis is an extinct genus of neoavian bird from the Early Eocene Fur Formation of Denmark. It contains a single named species, Morsoravis sedilis. Fossils of Morsoravis have also been found in the Green River Formation of Wyoming and possibly the Nanjemoy Formation of Virginia.
Scaniacypselus is an extinct genus of basal swifts from the Eocene of Denmark, Germany and France. Many well preserved specimens still showing plumage were discovered in the Messel Pit near Darmstadt, Germany. Species of Scaniacypselus had relatively broader wings than modern swifts and hindlimbs better adapted to perching on tree-branches, indicating that the bird was not as aerial as its extant relatives and likely nested in trees like hummingbirds and treeswifts. Two species are recognized, S. wardi and S. szarskii.
Tonsala is an extinct genus of Plotopteridae, a family of flightless seabird similar in biology with penguins, but more closely related to modern cormorants. The genus is known from terrains dated from the Late Oligocene of the State of Washington and Japan.
Lutavis is an extinct genus of potentially afroavian bird from the Early Eocene London Clay Formation of Essex, United Kingdom. The genus contains a single species, L. platypelvis, known from a partial skeleton.
Waltonavis is an extinct genus of potentially leptosomiform bird from the Early Eocene London Clay Formation of Essex, United Kingdom. The genus contains two species: W. paraleptosomus and W. danielsi, both known from partial skeletons.
Cyrilavis is an extinct genus of halcyornithid bird from the Early Eocene Fossil Butte Member of the Green River Formation, United States. The genus contains two species, Cyrilavis olsoni and Cyrilavis colburnorum.
Pulchrapollia is an extinct genus of halcyornithid bird from the Early Eocene London Clay of Walton-on-the-Naze, United Kingdom and the Nanjemoy Formation of Virginia, United States. The genus contains three species, Pulchrapollia gracilis, Pulchrapollia tenuipes and Pulchrapollia eximia.
Pseudasturides is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany, and possibly the Isle of Sheppey, United Kingdom. The genus is represented by a single species, Pseudasturides macrocephalus.
Serudaptus is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany. The genus contains one species, Serudaptus pohli, and is known for long, raptorial claws on its zygodactyl feet.
Parapsittacopes is an extinct genus of psittacopedid bird from the Early Eocene London Clay, United Kingdom. The genus contains one species, Parapsittacopes bergdahli.
Eofringillirostrum is an extinct species of bird known from the Early Eocene Green River Formation of the Western United States and from the Messel Pit in Germany. The genus contains two species, Eofringillirostrum boudreauxi and Eofringillirostrum parvulum. They are the earliest known fossil birds to have a finch-like beak.
Sororavis is an extinct genus of bird from the Early Eocene London Clay of Walton-on-the-Naze, United Kingdom. It contains a single named species, Sororavis solitarius. It is a member of the Morsoravidae.
Morsoravidae is a family of extinct birds known from the Early Eocene of Europe and North America, related to the Psittacopasseres, the clade containing parrots and songbirds. They have specialised feeding ecologies, and one morsoravid, Pumiliornis tesselatus, fed on nectar. The family contains the genera Morsoravis, Pumiliornis, and Sororavis.