Elektrithone Temporal range: | |
---|---|
Holotype forewing, Baltic amber | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Neuroptera |
Family: | Ithonidae |
Genus: | † Elektrithone Makarkin, Wedmann, & Weiterschan, 2014 |
Species: | †E. expectata |
Binomial name | |
†Elektrithone expectata Makarkin, Wedmann, & Weiterschan, 2014 | |
Elektrithone is an extinct genus of lacewing in the moth lacewings family Ithonidae. The genus is solely known from an Eocene fossil forewing found in Europe. At the times of description the genus was composed of a single species, Elektrithone expectata. [1]
Elektrithone expectata is known only from the one fossil, the holotype, specimen number SMF Be 2374, which is housed in the collections of the Senckenberg Forschungsinstitut und Naturmuseum in Frankfurt, Germany. The specimen is preserved as an inclusion in a transparent chunk of 46 million year old Baltic amber which was deposited during the Lutetian stage of the Middle Eocene. There is debate on what plant family produced the amber, with evidence supporting relatives of either an Agathis relative or a Pseudolarix relative. [2] The fossil was first studied by the paleoentomologists Vladimir N. Makarkin of the Far Eastern Branch of the Russian Academy of Sciences, Sonja Wedmann of the Senckenberg Forschungsstation, and Thomas Weiterschan of Höchst im Odenwald, Germany. Their 2014 type description of the new genus and species was published in the online journal Zootaxa . The genus name Elektrithone was coined by the researchers as a combination of the moth lacewing genus Ithone and the Greek word elektro meaning "amber", which is in reference to the finding of the genus in Baltic amber. [1] The specific epithet expectata is from the Latin word meaning "expected", alluding to the anticipated find of the family in Baltic amber. Elektrithone is the first member of Ithonidae to be discovered in Baltic amber. [1]
Using the Comstock–Needham system for describing insect venation patterns, the Elektrithone CuP (cubitus posterior, the "rear elbow") venation is similar to members of the neuropteran families Brongniartiellidae and Osmylopsychopidae, in that the CuP in both families is short and not comb like. However, both families have wings with an anal area that has at least one or two highly branched anal veins, a character that is not seen in Elektrithone. [1] The broad costal area of Elektrithone along with the strongly curved humeral veinlet, comb like MP (medial posterior) and CuA (cubitus anterior) veins and the series structure of the radial crossveins are all features seen in Ithonidae. The features found in Elektrithone are a mosaic of characters found in several genera of Ithonids. [1]
The holotype is composed of a single partially complete forewing which is missing a section of the central wing area. The preserved section of the forewing is approximately 19 mm (0.75 in) long and 10 mm (0.39 in) wide, with an estimated length, if whole, of 30 mm (1.2 in). The wing has a costal space that is wide, widening towards the wing base and a number of trichosors are present along the leading edge of the wing. [1] The ScA (subcostal anterior) veinlet is well developed, thick, and connecting to the ScP (subcostal posterior) vein before the humeral veinlet. The subcostal veinlets mostly are only shallowly forked, and the radial crossveins being grouped into a pair of gradate series. The wing does not show any evidence that nygma were present in life. [1]
Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments, and the two pairs are often referred to as the forewings and hindwings, respectively, though a few insects lack hindwings, even rudiments. The wings are strengthened by a number of longitudinal veins, which often have cross-connections that form closed "cells" in the membrane. The patterns resulting from the fusion and cross-connection of the wing veins are often diagnostic for different evolutionary lineages and can be used for identification to the family or even genus level in many orders of insects.
Hemerobiidae is a family of Neuropteran insects commonly known as brown lacewings, comprising about 500 species in 28 genera. Most are yellow to dark brown, but some species are green. They are small; most have forewings 4–10 mm long. These insects differ from the somewhat similar Chrysopidae not only by the usual coloring but also by the wing venation: hemerobiids differ from chrysopids in having numerous long veins and forked costal cross veins. Some genera are widespread, but most are restricted to a single biogeographical realm. Some species have reduced wings to the degree that they are flightless. Imagines (adults) of subfamily Drepanepteryginae mimic dead leaves. Hemerobiid larvae are usually less hairy than chrysopid larvae.
Psychopsidae is a family of winged insects of the order Neuroptera. They are commonly called silky lacewings.
Sisyridae, commonly known as spongeflies or spongillaflies, are a family of winged insects in the order Neuroptera. There are approximately 60 living species described, and several extinct species identified from the fossil record.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1878.
Microberotha is an extinct monotypic genus of "beaded lacewing" in the family Berothidae known from a fossil found in North America. When described the genus contained a single Ypresian-age species Microberotha macculloughi.
Allorapisma is an extinct genus of lacewing in the moth lacewings family Ithonidae. The genus is solely known from two Eocene fossils found in North America. At the time of description the genus was composed of a single species, Allorapisma chuorum.
Ithonidae, commonly called moth lacewings and giant lacewings, is a small family of winged insects of the insect order Neuroptera. The family contains a total of ten living genera, and over a dozen extinct genera described from fossils. The modern Ithonids have a notably disjunct distribution, while the extinct genera had a more global range. The family is considered one of the most primitive living neuropteran families. The family has been expanded twice, first to include the genus Rapisma, formerly placed in the monotypic family Rapismatidae, and then in 2010 to include the genera that had been placed into the family Polystoechotidae. Both Rapismatidae and Polystoechotidae have been shown to nest into Ithonidae sensu lato. The larvae of ithonids are grub-like, subterranean and likely phytophagous.
The Comstock–Needham system is a naming system for insect wing veins, devised by John Comstock and George Needham in 1898. It was an important step in showing the homology of all insect wings. This system was based on Needham's pretracheation theory that was later discredited by Frederic Charles Fraser in 1938.
Undulopsychopsis is an extinct genus of lacewing in the silky lacewing family Psychopsidae. The genus is solely known from a Cretaceous fossil found in China. Currently the genus is composed of a single species, Undulopsychopsis alexi.
Ainigmapsychops is an extinct genus of lacewing in the silky lacewing family Psychopsidae. The genus is solely known from an Eocene fossil found in North America. At the time of its description the new genus was composed of a single species, Ainigmapsychops inexspectatus.
Principiala is an extinct genus of lacewing in the moth lacewings family Ithonidae. The genus is known from Cretaceous fossils found in South America, Europe, and possibly Asia. The genus is composed of two species, the type species Principiala incerta, and Principiala rudgwickensis.
Wesmaelius mathewesi is an extinct species of lacewing in the neuropteran family Hemerobiidae known from an Eocene fossil found in North America
Cretomerobius is an extinct genus of lacewings in the neuropteran family Hemerobiidae known from fossils found in Asia. The genus currently contains a single species, the Aptian C. disjunctus.
Makarkinia is an extinct genus of lacewings in the family Kalligrammatidae described by Martins-Neto in 1997 from fossils found in the Crato Formation of the Araripe Basin in northeastern Brazil. The genus contains three species dating to the Late Aptian, Makarkinia adamsi, Makarkinia kerneri and Makarkinia irmae.
Proneuronema is an extinct genus of lacewings in the neuropteran family Hemerobiidae known from fossils found in amber. The genus currently contains three species, P. gradatum and P. minor from Baltic amber and P. sidorchukae from Rovno amber. A Ypresian fossil from Washington state was initially also placed within the genus, having been moved from its original placement as Cretomerobius wehri, but was subsequently moved to the separate genus Archibaldia by Vladimir Makarkin (2023).
Palaeopsychops is an extinct genus of lacewing in the moth lacewings family Ithonidae. The genus is known from Early Eocene fossils found in Europe, and North America and is composed of ten species. The ten species can be informally separated into two species groups based on veination of the forewings, the "European" and "North American" groups. When first described, the genus was placed in the family Psychopsidae, but later was moved to Polystoechotidae, which itself is now considered a subgroup of the moth lacewings.
The Prussian Formation, previously known as the Amber Formation, is a geologic formation in Prussia, today mostly Kaliningrad Oblast that dates to the Eocene. It holds 90% of the world's amber supply and Baltic amber is found exclusively in the Prussian Formation.
Polystoechotites is an extinct parataxon of lacewings in the moth lacewing family Ithonidae. The taxon is a collective group for fossil polystechotid giant lacewing species whose genus affiliation is uncertain, but which are distinct enough to identify as segregate species. Polystoechotites species are known from Eocene fossils found in North America and is composed of four named species Polystoechotites barksdalae, Polystoechotites falcatus, Polystoechotites lewisi, and Polystoechotites piperatus, plus two unnamed species. Three of the described species are known from fossils recovered from the Eocene Okanagan Highlands of Washington State, while the fourth is from Colorado.
Republica is an extinct zygopteran genus in the damselfly family Euphaeidae with a single described species, Republica weatbrooki. The species is solely known from the Early Eocene sediments exposed in the northeast of the U.S. state of Washington.