Equus neogeus

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Equus neogeus
Temporal range: Pleistocene (Ensenadan-Lujanian) ~1–0.012  Ma
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Amerhippus.jpg
Skull
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Genus: Equus
Species:
E. neogeus
Binomial name
Equus neogeus
Lund, 1840
Synonyms
  • Equus (Amerhippus) andium
  • Equus (Amerhippus) insulatus
  • Equus (Amerhippus) lasallei
  • Equus (Amerhippus) santaeelenae

Equus neogeus is an extinct species of equine native to South America during the Pleistocene. It was formerly thought to be several distinct species within the subgenus Amerhippus, but was later shown to be a single morphologically variable species. It is thought to be closely related to true horses.

Contents

Taxonomy

While they have formerly been referred to as belonging to 5 separate species, this has been revised down into three, [1] and more recently a single, morphologically variable species Equus neogeus. [2] [3] Some authors continue to recognise three species, restricting Equus neogeus to large-sized individuals spanning from the Pampas to Northeast Brazil, while separating the smaller Equus andium for populations in the northern-central Andes, and Equus insulatus for medium-sized animals spanning from Bolivia to Venezuela. These authors suggest that E. insulatus was the ancestor of the other species. [4] Historically, South American Equus species were placed in the subgenus Amerhippus, but this has subsequently been questioned. [4] A 2008 study of mitochondrial DNA fragments of a specimen of E. neogeus found it to be nested within mitochondrial lineages of E. caballus, [5] however, later studies suggested that this result required more specimens to be analysed for confirmation. [6] A close relationship to caballine horses was also supported by a 2019 morphological analysis study. [7]

Description

Life restoration Amerhippus A6 digital.jpg
Life restoration

Equus neogeus measured roughly 1.5 m (4.9 ft) tall and weighed approximately 400 kg (880 lb). [8]

Distribution

They were one of two groups of equines in South America, alongside Hippidion . [9] Fossils have been recovered from the Tarija Formation of Bolivia, the Serranía del Perijá in Venezuela, the Chiu-Chiu Formation of Chile, the Sabana Formation of the Bogotá savanna in Colombia, [10] and from various locations in Ecuador. [8] Equus first appeared in South America during the late Early Pleistocene-earliest Middle Pleistocene, around 1 to 0.8 million years ago, based on remains found near Tarija, Bolivia. [11]

Paleobiology

A 2019 study suggested that Equus neogeus specimens from Uruguay were primarily grazers that fed on both C4 and C3 grasses in prairies and open woodlands. [12]

Extinction

Equus neogeus became extinct at the end of the Late Pleistocene as part of the end-Pleistocene extinctions, along with the vast majority of other large mammals in South America. [13] The youngest remains date to approximately 11,700 years Before Present (BP), in Río Quequén Salado, in the southwest of Buenos Aires province, Argentina. [11] The extinctions followed human arrival to the Americas, and several sites show evidence of human interaction with Equus neogeus, which may have been a factor in its extinction. [14] At the Paso Otero 5 site in the Pampas of northeast Argentina, Fishtail points are associated with burned bones of Equus neogeus and other extinct megafauna. The bones appear to have been deliberately burned as a source of fuel. Due to the poor preservation of the bones there is no clear evidence of human modification. [15] At the Taguatagua 3 in central Chile, dating to around 12,440–12,550 years BP, a first phalanx of Equus was found near a hearth, alongside the remains of other megafuna, including the much more abundant remains of gomphotheres (elephant relatives). [16] At the Arroyo Seco 2 site in the Argentinian Pampas, which has multiple episodes of human activity dating to between 14,782 and 11,142 cal yr BP remains of Equus neogeus (alongside those of other megafauna) are associated with human artifacts and exhibit fracture marks indicative of butchery. [14] [17]

Related Research Articles

<span class="mw-page-title-main">Equidae</span> Family of hoofed mammals

Equidae is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. The family evolved around 50 million years ago from a small, multi-toed ungulate into larger, single-toed animals. All extant species are in the genus Equus, which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families. It is more specifically grouped within the superfamily Equoidea, the only other family being the extinct Palaeotheriidae.

<i>Megatherium</i> Genus of ground sloth (extinct)

Megatherium is an extinct genus of ground sloths endemic to South America that lived from the Early Pliocene through the end of the Late Pleistocene. It is best known for the elephant-sized type species Megatherium americanum, native to northern Patagonia and the Pampas northwards to southern Bolivia during the late Middle Pleistocene and Late Pleistocene. Various other species belonging to the subgenus Pseudomegatherium ranging in size comparable to considerably smaller than M. americanum are known from the Andean region.

<span class="mw-page-title-main">Megafauna</span> Large animals

In zoology, megafauna are large animals. The precise definition of the term varies widely, though a common threshold is approximately 45 kilograms (99 lb), with other thresholds as low as 10 kilograms (22 lb) or as high as 1,000 kilograms (2,200 lb). Large body size is generally associated with other traits, such as having a slow rate of reproduction and in large herbivores, reduced or negligible adult mortality from being killed by predators.

<span class="mw-page-title-main">Gomphothere</span> Extinct family of proboscidean mammals

Gomphotheres are an extinct group of proboscideans related to modern elephants. First appearing in Africa during the Oligocene, they dispersed into Eurasia and North America during the Miocene and arrived in South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae.

<span class="mw-page-title-main">Late Pleistocene</span> Third division (unofficial) of the Pleistocene Epoch

The Late Pleistocene is an unofficial age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a stratigraphic perspective. It is intended to be the fourth division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently defined as the time between c. 129,000 and c. 11,700 years ago. The late Pleistocene equates to the proposed Tarantian Age of the geologic time scale, preceded by the officially ratified Chibanian. The beginning of the Late Pleistocene is the transition between the end of the Penultimate Glacial Period and the beginning of the Last Interglacial around 130,000 years ago. The Late Pleistocene ends with the termination of the Younger Dryas, some 11,700 years ago when the Holocene Epoch began.

<i>Macrauchenia</i> Extinct genus of camel-like ungulate

Macrauchenia is an extinct genus of large ungulate native to South America from the Pliocene or Middle Pleistocene to the end of the Late Pleistocene. It is a member of the extinct order Litopterna, a group of South American native ungulates distinct from the two orders which contain all living ungulates which had been present in South America since the early Cenozoic, over 60 million years ago, prior to the arrival of living ungulates in South America around 2.5 million years ago as part of the Great American Interchange. The bodyform of Macrauchenia has been described as similar to a camel, being one of the largest-known litopterns, with an estimated body mass of around 1 tonne. The genus gives its name to its family, Macraucheniidae, which like Macrauchenia typically had long necks and three-toed feet, as well as a retracted nasal region, which in Macrauchenia manifests as the nasal opening being on the top of the skull between the eye sockets. This has historically been argued to correspond to the presence of a tapir-like proboscis, though recent authors suggest a moose-like prehensile lip or a saiga antelope-like nose to filter dust are more likely.

<i>Equus scotti</i> Extinct species of mammal

Equus scotti is an extinct species of horse native to Pleistocene North America.

<span class="mw-page-title-main">Wild horse</span> Undomesticated four-footed mammal from the equine family

The wild horse is a species of the genus Equus, which includes as subspecies the modern domesticated horse as well as the endangered Przewalski's horse. The European wild horse, also known as the tarpan, that went extinct in the late 19th or early 20th century has previously been treated as the nominate subspecies of wild horse, Equus ferus ferus, but more recent studies have cast doubt on whether tarpans were truly wild or if they actually were feral horses or hybrids.

<i>Cuvieronius</i> Extinct genus of mammals

Cuvieronius is an extinct New World genus of gomphothere which ranged from southern North America to western South America during the Pleistocene epoch. Among the last gomphotheres, it became extinct at the end of the Pleistocene, approximately 12,000 years ago, following the arrival of humans to the Americas.

<i>Equus namadicus</i> Extinct species of mammal

Equus namadicus is a prehistoric equid, known from remains dating to the Middle and Late Pleistocene from across the Indian subcontinent, with its last dated records being approximately 29-14,000 years ago. It is considered a "stenonine horse", meaning that it is probably more closely related to zebras and asses than true horses. It is relatively large in size. It is very similar to the earlier Equus sivalensis, also from the Indian subcontinent, from which it only differs in size and in subtle aspects of dental anatomy, and it has sometimes been suggested to be a synonym of it.

<i>Hippidion</i> Extinct genus of mammals

Hippidion is an extinct genus of equine that lived in South America from the Late Pliocene to the end of the Late Pleistocene (Lujanian), between 2.5 million and 11,000 years ago. They were one of two lineages of equines native to South America during the Pleistocene epoch, alongside Equus (Amerhippus) neogeus.

<span class="mw-page-title-main">Evolution of the horse</span>

The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling Eohippus into the modern horse. Paleozoologists have been able to piece together a more complete outline of the evolutionary lineage of the modern horse than of any other animal. Much of this evolution took place in North America, where horses originated but became extinct about 10,000 years ago, before being reintroduced in the 15th century.

Plesippus is a genus of extinct horse from the Pleistocene of North America. Although commonly seen as a subgenus of Equus recent cladistic analysis considers it a distinct genus.

<span class="mw-page-title-main">Hagerman horse</span> Extinct species of mammal

Equus simplicidens, sometimes known as the Hagerman horse or the American zebra is an extinct species in the horse family native to North America during the Pliocene and Early Pleistocene. It is one of the oldest and most primitive members of the genus Equus. It is the state fossil of Idaho, where its abundant remains were discovered in 1928, specifically in Hagerman, Idaho. They are also known from other US states including Arizona, California, Texas, Nebraska and Kansas.

<span class="mw-page-title-main">Late Pleistocene extinctions</span> Extinctions of large mammals in the Late Pleistocene

The Late Pleistocene to the beginning of the Holocene saw the extinction of the majority of the world's megafauna, which resulted in a collapse in faunal density and diversity across the globe. The extinctions during the Late Pleistocene are differentiated from previous extinctions by its extreme size bias towards large animals, and widespread absence of ecological succession to replace these extinct megafaunal species, and the regime shift of previously established faunal relationships and habitats as a consequence. The timing and severity of the extinctions varied by region and are thought to have been driven by varying combinations of human and climatic factors. Human impact on megafauna populations is thought to have been driven by hunting ("overkill"), as well as possibly environmental alteration. The relative importance of human vs climatic factors in the extinctions has been the subject of long-running controversy.

<i>Glyptodon</i> Genus of large, heavily armored mammals

Glyptodon is a genus of glyptodont, an extinct group of large, herbivorous armadillos, that lived from the Pliocene, around 3.2 million years ago, to the early Holocene, around 11,000 years ago, in South America. It is one of, if not the, best known genus of glyptodont. Glyptodon has a long and storied past, being the first named extinct cingulate and the type genus of the subfamily Glyptodontinae. Fossils of Glyptodon have been recorded as early as 1814 from Pleistocene aged deposits from Uruguay, though many were incorrectly referred to the ground sloth Megatherium by early paleontologists.

<i>Haringtonhippus</i> Extinct genus of mammals

Haringtonhippus is an extinct genus of equine from the Pleistocene of North America The genus is monospecific, consisting of the species H. francisci, initially described in 1915 by Oliver Perry Hay as Equus francisci. Members of the genus are often referred to as stilt-legged horses, in reference to their slender distal limb bones, in contrast with those of contemporary "stout legged" caballine true horses.

<i>Eutatus</i> Extinct genus of mammals

Eutatus is an extinct genus of large armadillos of the family Chlamyphoridae. It was endemic to South America from the Early Miocene to Late Pleistocene, living from 17.5 Ma-11,000 years ago, with possible survival into the early Holocene and existing for approximately 17.49 million years. Based on carbon isotope ratios, it is thought to have been an herbivore that fed on grasses.

<i>Notiomastodon</i> Extinct genus of gomphothere elephantimorph native to South America

Notiomastodon is an extinct genus of gomphothere proboscidean, endemic to South America from the Pleistocene to the beginning of the Holocene. Notiomastodon specimens reached a size similar to that of the modern Asian elephant, with a body mass of 3-4 tonnes. Like other brevirostrine gomphotheres such as Cuvieronius and Stegomastodon, Notiomastodon had a shortened lower jaw and lacked lower tusks, unlike more primitive gomphotheres like Gomphotherium.

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