Equus neogeus

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Equus neogeus
Temporal range: Pleistocene (Ensenadan-Lujanian) ~1–0.012  Ma
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Amerhippus.jpg
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Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Genus: Equus
Species:
E. neogeus
Binomial name
Equus neogeus
Lund, 1840
Synonyms
  • Equus (Amerhippus) andium
  • Equus (Amerhippus) insulatus
  • Equus (Amerhippus) lasallei
  • Equus (Amerhippus) santaeelenae

Equus neogeus is an extinct species of equine native to South America during the Pleistocene. It was formerly thought to be several distinct species within the subgenus Amerhippus, but was later shown to be a single morphologically variable species. It is thought to be closely related to true horses.

Contents

Taxonomy

While they have formerly been referred to as belonging to 5 separate species, this has been revised down into three, [1] and more recently a single, morphologically variable species Equus neogeus. [2] [3] Some authors continue to recognise three species, restricting Equus neogeus to large-sized individuals spanning from the Pampas to Northeast Brazil, while separating the smaller Equus andium for populations in the northern-central Andes, and Equus insulatus for medium-sized animals spanning from Bolivia to Venezuela. These authors suggest that E. insulatus was the ancestor of the other species. [4] Historically, South American Equus species were placed in the subgenus Amerhippus, but this has subsequently been questioned. [4] A 2008 study of mitochondrial DNA fragments of a specimen of E. neogeus found it to be nested within mitochondrial lineages of E. caballus, [5] however, later studies suggested that this result required more specimens to be analysed for confirmation. [6] A close relationship to caballine horses was also supported by a 2019 morphological analysis study. [7]

Description

Life restoration Amerhippus A6 digital.jpg
Life restoration

Equus neogeus measured roughly 1.5 m (4.9 ft) tall and weighed approximately 400 kg (880 lb). [8]

Distribution

They were one of two groups of equines in South America, alongside Hippidion . [9] Fossils have been recovered from the Tarija Formation of Bolivia, the Serranía del Perijá in Venezuela, the Chiu-Chiu Formation of Chile, the Sabana Formation of the Bogotá savanna in Colombia, [10] and from various locations in Ecuador. [8] Equus first appeared in South America during the late Early Pleistocene-earliest Middle Pleistocene, around 1 to 0.8 million years ago, based on remains found near Tarija, Bolivia. [11]

Paleobiology

A 2019 study suggested that Equus neogeus specimens from Uruguay were primarily grazers that fed on both C4 and C3 grasses in prairies and open woodlands. [12]

Extinction

Equus neogeus became extinct at the end of the Late Pleistocene as part of the Quaternary extinction event, along with the vast majority of other large mammals in South America. [13] The youngest remains date to approximately 11,700 years Before Present (BP), in Río Quequén Salado, in the southwest of Buenos Aires province, Argentina. [11] At the Arroyo Seco 2 site in the Argentinian Pampas, remains of Equus neogeus are associated with human artifacts, indicating that they were butchered by humans at the site, which dates to 14,782–11,142 cal yr BP, suggesting that hunting by recently arrived humans may have played a role in its extinction. [14]

Related Research Articles

<span class="mw-page-title-main">Equidae</span> Family of hoofed mammals

Equidae is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. The family evolved around 50 million years ago from a small, multi-toed ungulate into larger, single-toed animals. All extant species are in the genus Equus, which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families.

<span class="mw-page-title-main">Proboscidea</span> Order of elephant-like mammals

The Proboscidea are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. From the mid-Miocene onwards, most proboscideans were very large. The largest land mammal of all time may have been a proboscidean; Palaeoloxodon namadicus was up to 5.2 m (17.1 ft) at the shoulder and may have weighed up to 22 t, almost double the weight of some sauropods like Diplodocus carnegii. The largest extant proboscidean is the African bush elephant, with a record of size of 4 m (13.1 ft) at the shoulder and 10.4 t. In addition to their enormous size, later proboscideans are distinguished by tusks and long, muscular trunks, which were less developed or absent in early proboscideans.

<span class="mw-page-title-main">Mastodon</span> Genus of mammals (fossil)

A mastodon is any proboscidean belonging to the extinct genus Mammut. Mastodons inhabited North and Central America from the late Miocene up to their extinction at the end of the Pleistocene 10,000 to 11,000 years ago.

<i>Equus</i> (genus) Genus of mammals which includes horses, donkeys, and zebras

Equus is a genus of mammals in the family Equidae, which includes horses, asses, and zebras. Within the Equidae, Equus is the only recognized extant genus, comprising seven living species. Like Equidae more broadly, Equus has numerous extinct species known only from fossils. The genus most likely originated in North America and spread quickly to the Old World. Equines are odd-toed ungulates with slender legs, long heads, relatively long necks, manes, and long tails. All species are herbivorous, and mostly grazers, with simpler digestive systems than ruminants but able to subsist on lower-quality vegetation.

<span class="mw-page-title-main">Gomphothere</span> Extinct family of proboscidean mammals

Gomphotheres are an extinct group of proboscideans related to modern elephants. They were widespread across Afro-Eurasia and North America during the Miocene and Pliocene epochs and dispersed into South America during the Pleistocene as part of the Great American Interchange. Gomphotheres are a paraphyletic group that is ancestral to Elephantidae, which contains modern elephants, as well as Stegodontidae. While most famous forms such as Gomphotherium had long lower jaws with tusks, which is the ancestral condition for the group, some later members developed shortened (brevirostrine) lower jaws with either vestigial or no lower tusks, looking very similar to modern elephants, an example of parallel evolution, which outlasted the long-jawed gomphotheres. By the end of the Early Pleistocene, gomphotheres became extinct in Afro-Eurasia, with the last two genera, Cuvieronius ranging from southern North America to eastern South America, and Notiomastodon having a wide range over most of South America until the end of the Pleistocene around 12,000 years ago, when they became extinct following the arrival of humans.

<span class="mw-page-title-main">Wild horse</span> Undomesticated four-footed mammal from the equine family

The wild horse is a species of the genus Equus, which includes as subspecies the modern domesticated horse as well as the endangered Przewalski's horse. The European wild horse, also known as the tarpan, that went extinct in the late 19th or early 20th century has previously been treated as the nominate subspecies of wild horse, Equus ferus ferus, but more recent studies have cast doubt on whether tarpans were truly wild or if they actually were feral horses or hybrids.

<span class="mw-page-title-main">European wild ass</span> Extinct species of mammal

The European wild ass or hydruntine is an extinct equine from the Middle Pleistocene to Late Holocene of Europe and West Asia, and possibly North Africa. It is a member of the subgenus Asinus, and closely related to the living Asiatic wild ass. The specific epithet, hydruntinus, means from Otranto.

<i>Cuvieronius</i> Extinct genus of mammals

Cuvieronius is an extinct New World genus of gomphothere which ranged from southern North America to eastern South America during the Pleistocene epoch. Among the last gomphotheres, it became extinct at the end of the Pleistocene, approximately 12,000 years ago, following the arrival of humans to the Americas.

<i>Stegomastodon</i> Extinct genus of mammals

Stegomastodon is an extinct genus of gomphotheres, a family of proboscideans. It ranged throughout North America from the early Blancan ~4 Ma, to the early Irvingtonian. The South American species have been synonymized with Notiomastodon platensis.

<i>Equus sivalensis</i> Extinct species of mammal

Equus sivalensis is an extinct species of large equid native to the northern Indian subcontinent. Remains date from the beginning of the Pleistocene, c. 2.58 million years ago until around 600,000 years ago, during the Middle Pleistocene. It is considered a "stenonine horse", meaning that it is more closely related to zebras and asses than true horses. Based on isotopes and teeth morphology, it is thought to have been a grazer. The later species Equus namadicus from the same region has sometimes been suggested to be a synonym due to their similar teeth morphology.

<i>Equus namadicus</i> Extinct species of mammal

Equus namadicus is a prehistoric equid, known from remains dating to the Middle and Late Pleistocene from across the Indian subcontinent, with its last dated records being approximately 29-14,000 years ago. It is considered a "stenonine horse", meaning that it is probably more closely related to zebras and asses than true horses. It is relatively large in size. It is very similar to the earlier Equus sivalensis, also from the Indian subcontinent, from which it only differs in size and in subtle aspects of dental anatomy, and it has sometimes been suggested to be a synonym of it.

<i>Hippidion</i> Extinct genus of mammals

Hippidion is an extinct genus of equine that lived in South America from the Late Pliocene to the end of the Late Pleistocene (Lujanian), between 2.5 million and 11,000 years ago. They were one of two lineages of equines native to South America during the Pleistocene epoch, alongside Equus (Amerhippus) neogeus.

<span class="mw-page-title-main">Evolution of the horse</span> Derivation of horses from an ungulate precursor

The evolution of the horse, a mammal of the family Equidae, occurred over a geologic time scale of 50 million years, transforming the small, dog-sized, forest-dwelling Eohippus into the modern horse. Paleozoologists have been able to piece together a more complete outline of the evolutionary lineage of the modern horse than of any other animal. Much of this evolution took place in North America, where horses originated but became extinct about 10,000 years ago, before being reintroduced in the 15th century.

Plesippus is a genus of extinct horse from the Pleistocene of North America. Although commonly seen as a subgenus of Equus recent cladistic analysis considers it a distinct genus.

<i>Haringtonhippus</i> Extinct genus of mammals

Haringtonhippus is an extinct genus of equine from the Pleistocene of North America The genus is monospecific, consisting of the species H. francisci, initially described in 1915 by Oliver Perry Hay as Equus francisci. Prior to the naming of the genus, it was sometimes referred to as the New World stilt-legged horse, in reference to its slender distal limb bones, in contrast with those of contemporary "stout legged" caballine true horses.

<i>Notiomastodon</i> Extinct genus of mammals

Notiomastodon is an extinct genus of gomphothere proboscidean, endemic to South America from the Pleistocene to the beginning of the Holocene. Notiomastodon specimens reached a size similar to that of the modern Asian elephant. Like other brevirostrine gomphotheres such as Cuvieronius and Stegomastodon, Notiomastodon had a shortened lower jaw and lacked lower tusks.

<i>Arctotherium</i> Extinct genus of bears

Arctotherium is an extinct genus of the Pleistocene short-faced bears endemic to Central and South America. Arctotherium migrated from North America to South America during the Great American Interchange, following the formation of the Isthmus of Panama during the late Pliocene. The genus consists of one early giant form, A. angustidens, and several succeeding smaller species, which were within the size range of modern bears. Arctotherium was adapted to open and mixed habitat. They are genetically closer to the spectacled bear, than to Arctodus of North America, implying the two extinct forms evolved large size in a convergent manner.

Equus alaskae was a Pleistocene species of horse, now extinct, that inhabited North America.

<i>Equus stenonis</i> Extinct species of mammal

Equus stenonis is an extinct species of equine that lived in Western Eurasia during the Early Pleistocene epoch.

References

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