Eschrichtioides Temporal range: | |
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Eschrichtioides gastaldii, from Portis 1885 | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Infraorder: | Cetacea |
Family: | Eschrichtiidae |
Genus: | † Eschrichtioides Bisconti 2008 |
Species | |
†E. gastaldii | |
Synonyms | |
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Eschrichtioides is an extinct genus of baleen whale known from the early Pliocene of Northern Italy. Its type species, E. gastaldii, had a complex taxonomic history, starting as a cetothere, then as an extinct member of Balaenoptera , before being finally recognized as a relative of the gray whale.
Eschrichtioides is one of two Eschrichtius relatives known from the Neogene of Italy, the other being Archaeschrichtius . Its holotype, MRSN 13802, comes from the early Pliocene-age Sabbie d'Asti Formation of the Piedmont region in Italy [1] and it is currently exposed in Asti's paleontological museum "Museo Paleontologico Territoriale dell'Astigiano". [2]
Like other cetaceans from the Italian Pliocene, the classification of Eschrichtioides was highly contentious.
Brandt 1873 [3] described several species of Cetotherium from the Pliocene of northern Italy, including C. cortesi based on a specimen (MSRN 13802) found by Italian geologist Giuseppe Cortesi in 1816 near the town Cortandone ( 45°00′N8°06′E / 45.0°N 8.1°E : paleocoordinates 44°54′N8°00′E / 44.9°N 8.0°E ) [4]
Cortesi 1819 [5] described how he, after a day of searching for fossils, found what he first thought was a small piece of wood in a small river. After a closer inspection it proved to be a bone, and Cortesi and his companion found more and more of them in the banks of the river. After four days of digging they had unveiled a giant skeleton easily identifiable as a whale 12 feet 5 inches long. The three feet long head and its dentaries had no signs of either teeth nor tooth sockets. Cortesi noted that at that time few naturalists could assign cetacean fossils to individual species, and he therefore never named his specimen.
Desmoulins 1822 [6] thought the "Baleine de Cortesi" represented a distinctive species because it was a very small adult individual and because the curvature of the maxillary branches was less convex than in any other known whale. Other French naturalists [7] named it "Le rorqual de Cortési"; arguing that both Cortesi's fossil whales ("rorqual de Cortési" and "de Cuvier") were closely related to extant rorquals and the only difference between them was the much smaller size of the former.
Brandt 1873 [8] assigned the "baleine de Cortesi" to his own genus Cetotherium Cortesii.
In a preliminary note Strobel 1875 [9] pointed out differences in cranial morphology in two skulls both described as "Cetotherium cortesii" (a species named and described by Brandt 1873), and he therefore proposed the new specific name "Cetotherium gastaldii" (in honour of Italian palaeontologist Bartolomeo Gastaldi) for one of the specimens. Strobel 1881 [10] described the specimen and reiterated his arguments from 1875.
Originally misidentified as the balaenopterid "Balaena" cortesii, Brandt 1873 described "B." cortesii as a species of Cetotherium. [8] Strobel 1875 finally named it C. gastaldii. [11] However, several authors considered it to be referrable to Balaenoptera , [12] a taxonomic arrangement which was accepted for more than a century. [13] [14] However, re-examination of MRSN 13802 revealed that "C." gastaldii was not a balaenopterid and instead constituted its own genus within Eschrichtiidae. [1]
Eschrichtiidae or the gray whales is a family of baleen whale with a single extant species, the gray whale, as well as four described fossil genera: Archaeschrichtius (Miocene), Glaucobalaena and Eschrichtioides (Pliocene) from Italy, and Gricetoides from the Pliocene of North Carolina. Some phylogenetic studies have found this family to be invalid, with its members nesting inside of the clade Balaenopteridae. The names of the extant genus and the family honours Danish zoologist Daniel Eschricht.
Neobalaenidae is a family of baleen whales including the extant pygmy right whale. Although traditionally considered related to balaenids, recent studies by Fordyce and Marx (2013) and Ludovic Dutoit and colleagues (2023) have recovered the living pygmy right whale as a member of Cetotheriidae, making it the only extant cetotheriid. Not all authors agree with this placement.
Balaenoptera is a genus of rorquals containing eight extant species. Balaenoptera comprises all but two of the extant species in its family ; the genus is currently polyphyletic, with the two aforementioned species being phylogenetically nested within it.
Cetotherium is an extinct genus of baleen whales from the family Cetotheriidae.
Parabalaenoptera is a genus of prehistoric baleen whale found in Marin County, California. The type species is P. baulinensis. It was estimated to be about the size of the modern gray whale, about 16 metres (52 ft) long. It lived during the late Miocene.
Plesiobalaenoptera is a genus of extinct rorqual which existed in Italy during the late Miocene epoch. The type species is P. quarantellii. It is the oldest known rorqual from the Mediterranean basin. Fossils have been found from sediments of the Stirone River in Northern Italy that were deposited during the Tortonian age, around 11 to 7 million years ago.
Diunatans is an extinct genus of rorqual. It lived in the North Sea during the Early Pliocene. Two specimens have been found from the Netherlands. They were collected from the Kattendijk Formation in the province of Zeeland, which is Zanclean in age. Diunatans is considered to be a stem balaenopterid because it falls outside the Balaenoptera+Megaptera clade, which includes all living rorquals.
Cetotheriidae is a family of baleen whales. The family is known to have existed from the Late Oligocene to the Early Pleistocene before going extinct. Although some phylogenetic studies conducted by Fordyce & Marx 2013 recovered the living pygmy right whale as a member of Cetotheriidae, making the pygmy right whale the only living cetotheriid, other authors either dispute this placement or recover Neobalaenidae as a sister group to Cetotheriidae.
Parietobalaena is an extinct genus of baleen whale, belonging to the family Pelocetidae. Fossils are found in Miocene-aged marine strata in North America, Europe, Australia, and Japan, including the Temblor and Itahashi formations. Based on previous estimates of juvenile specimens, Tsai (2017) suggested a body size of 12-15 m for P. yamaokai, akin to that of the gray whale.
Piscobalaena is an extinct genus of cetaceans, which lived from the Middle to Late Miocene epochs in Peru and Florida. Its fossils have been found in the Pisco Formation of Peru and the Bone Valley Formation of Florida. At least some individuals of this diminutive whale were preyed on by the shark O. megalodon.
Herpetocetus is a genus of cetotheriid mysticete in the subfamily Herpetocetinae. Considerably smaller than modern baleen whales, Herpetocetus measured only 3 to 4 meters in length. Additionally, due to the structure of its jaw, it was unable to open its mouth as wide as modern baleen whales, making it incapable of lunge feeding.
Protororqualus is a genus of extinct rorqual from the late Pliocene of Mount Pulgnasco, Italy.
Cetotheriophanes is an extinct rorqual from the late Pliocene (Piacenzian) of Northern Italy.
Archaebalaenoptera is a genus of extinct rorqual known from late Miocene to Pliocene-age marine deposits of the Netherlands, Northern Italy, and Peru.
Plesiocetus is a genus of extinct rorquals found worldwide. It has had a chequered taxonomic history, having served as a wastebasket genus for a handful of mysticete species.
Aglaocetus is a genus of extinct baleen whales known from the Miocene of Patagonia, the US Eastern Seaboard, Japan and the Low Countries. It was once considered a member of Cetotheriidae along with many other putative cetotheres, but was recently recognized as representing a distinct family from true Cetotheriidae.
"Balaenoptera" ryani is an extinct species of Balaenopteridae from the late Miocene of California. It was originally considered a species of Balaenoptera, but is now recognized as more primitive than any extant or fossil balaenopterid and thus in need of a new generic name.
Nannocetus is an extinct baleen whale belonging to the family Cetotheriidae.
Fragilicetus is an extinct genus of rorqual from Early Pliocene deposits in Belgium.