| Horopeta | |
|---|---|
 | |
| Restoration of Horopeta | |
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Artiodactyla |
| Infraorder: | Cetacea |
| Family: | incertae sedis |
| Genus: | † Horopeta Tsai and Fordyce, 2015 |
| Species | |
| |
Horopeta is a genus of baleen whale from the Late Oligocene (Chattian) Kokoamu Greensand of New Zealand.
Horopeta can be distinguished from other balaenomorphs in the following characters: base of frontal sloping moderately from skull midline; parietal considerably exposed at sagittal crest; parietal largely exposed at sagittal crest; unfused and short posterior process of periotic and tympanic bulla; prominent superior process of periotic; presence of elliptical foramen and sigmoidal cavity in tympanic bulla; unfused and short posterior processes of periotic and tympanic bulla; a distinct pedicle plate situated in fovea epitubaria; presence of horizontal cleft, elliptical foramen, and sigmoidal cavity in tympanic bulla. [1]
Initial cladistic analysis recovered Horopeta as either a basal balaenomorph or a basal cetotheriid, although the describers noted that the juvenile nature of the holotype might influence its cladistic position within Chaeomysticeti. Later cladistic analyses corroborated the placement of Horopeta outside crown Mysticeti. [1] [2] [3]
Based on the structure of the jaw, Horopeta was one of the earliest chaeomysticetes capable of gulp-feeding as in extant mysticetes. [1] The same deposits that yielded Horopeta have also yielded the primitive odontocetes Awamokoa , Austrosqualodon , Otekaikea , and Waipatia , the eomysticetids Matapanui , Tohoraata , Tokarahia , and Waharoa , and the balaenomorphs Mauicetus and Whakakai .
Neobalaenidae is a family of baleen whales including the extant pygmy right whale. Although traditionally considered related to balaenids, a recent phylogenetic study by Fordyce and Marx (2013) recovered the living pygmy right whale as a member of Cetotheriidae, making it the only extant cetotheriid, but not all authors agree with this argument.
Janjucetus is an extinct genus of cetacean, and a basal baleen whale (Mysticeti), from the Late Oligocene around 25 million years ago (mya) off southeast Australia, containing one species J. hunderi. Unlike modern mysticetes, it possessed large teeth for gripping and shredding prey, and lacked baleen, and so was likely to have been a predator that captured large single prey animals rather than filter feeding. However, its teeth may have interlocked, much like those of the modern-day filter-feeding crabeater seal, which would have allowed some filter-feeding behavior. Its hunting behaviour was probably similar to the modern-day leopard seal, probably eating large fish. Like baleen whales, Janjucetus could not echolocate; however, it did have unusually large eyes, and so probably had an acute sense of vision. The only specimen was found on the Jan Juc beach, where the remains of the extinct whales Mammalodon, Prosqualodon and Waipatia have also been discovered.
Mammalodon is an extinct genus of archaic baleen whale belonging to the family Mammalodontidae.
Mammalodontidae is a family of extinct whales known from the Oligocene of Australia and New Zealand.
Aetiocetus is a genus of extinct basal mysticete, or baleen whale that lived 33.9 to 23.03 million years ago, in the Oligocene in the North Pacific ocean, around Japan, Mexico, and Oregon, U.S. It was first described by Douglas Emlong in 1966 and currently contains known four species, A. cotylalveus, A. polydentatus, A. tomitai, and A. weltoni. These whales are remarkable for their retention of teeth and presence of nutrient foramina, indicating that they possessed baleen. Thus, Aetiocetus represents the transition from teeth to baleen in Oligocene mysticetes. Baleen is a highly derived character, or synapomorphy, of mysticetes, and is a keratinous structure that grows from the palate, or roof of the mouth, of the whale. The presence of baleen is inferred from the fossil record in the skull of Aetiocetus. Aetiocetus is known from both sides of the Pacific Ocean: it was first documented in Oregon, United States, but it is also known from Japan and Mexico. The genus is currently constrained to the Northern hemisphere and has little value in biostratigraphic studies of the Oligocene due to its limited occurrences across the Pacific.
The Kokoamu Greensand is a geological formation found in New Zealand. It is a fossil-bearing, late Oligocene, greensand rock unit of the eastern South Island, especially the Waitaki District of North Otago and the southern Canterbury region. The formation was named by geologist Maxwell Gage in the 1950s. In North Otago it underlies the thicker and harder Otekaike Limestone. The formation gets its green colour from the mineral glauconite which forms slowly on the ocean floor.
Eomysticetus is an extinct genus of baleen whale from the late Oligocene (Chattian) Chandler Bridge Formation of South Carolina.
Nannocetus is an extinct baleen whale belonging to the family Cetotheriidae.
Otekaikea is an extinct genus of toothed whale closely related to Waipatia. It is known from the late Oligocene (Chattian) of New Zealand.
Mauicetus is a genus of extinct baleen whale from the Late Oligocene of New Zealand.
Tohoraata is a genus of eomysticetid baleen whale from the Late Oligocene (Chattian) of New Zealand. There are two recognized species, T. raekohao and T. waitakiensis.
Eomysticetidae is a family of extinct mysticetes belonging to Chaeomysticeti. It is one of two families in the basal chaeomysticete clade Eomysticetoidea.
Tranatocetus is an extinct genus of mysticete from the late Miocene (Tortonian) of Jutland, Denmark. The type and only species is Tranatocetus argillarius.
Tokarahia is a genus of eomysticetid baleen whale from the Late Oligocene (Chattian) of New Zealand. There are two recognized species, T. kauaeroa and T. lophocephalus.
Aetiocetidae is an extinct family of toothed baleen whales known from the Oligocene. The whales are from the North Pacific Ocean and ranged in size from 3 to 8 metres long. Many of the described specimens were discovered from the Upper Oligocene of the Japanese Morawan Formation, the largest known one from the Morawan's Upper tuffaceous siltstone. Other formally described extinct toothed mysticetis from this time are smaller, from 3 to 4 metres in length. Mysticeti with true baleen are seen in fossils from the Upper Oligocene. The monophyly of the family is still uncertain, as are the evolutionary relationship between the early toothed baleen whales and the early and extant edentulous baleen whales. However, the cladistic analyses of Coronodon and Mystacodon seem to indicate that Aetiocetidae and Llanocetidae are more closely related to crown Mysticeti than to Mammalodontidae, Coronodon, and Mystacodon.
Waharoa is a genus of eomysticetid baleen whale from the Late Oligocene (Chattian) of New Zealand. It was identified with the discovery of Waharoa ruwhenua by Boessenecker and Fordyce (2015), which added a new genus and species to a monophyletic family Eomysticetidae.
Matapanui is a genus of eomysticetid baleen whale from the Late Oligocene Kokoamu Greensand of New Zealand.
Whakakai is a genus of baleen whale from the Late Oligocene (Chattian) Kokoamu Greensand of New Zealand.
Toipahautea is a genus of baleen whale from the Late Oligocene (Chattian) Kokoamu Greensand of New Zealand.
