European dhole | |
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Skeletal remains dating back to upper Würm period from Cova del Parpalló, Gandía, Valencia, Spain | |
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Life restoration | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Family: | Canidae |
Genus: | Cuon |
Species: | |
Subspecies: | †C. a. europaeus |
Trinomial name | |
†Cuon alpinus europaeus Bourguignat, 1868 |
The European dhole (Cuon alpinus europaeus) was a paleosubspecies of the dhole, which ranged throughout much of Western and Central Europe during the Middle and Late Pleistocene. Like the modern Asiatic populations, it was a more progressive form than other prehistoric members of the genus Cuon, having transformed its lower molar tooth into a single cusped slicer. It was virtually indistinguishable from its modern counterpart, save for its greater size, which closely approached that of the gray wolf. [1]
Cuon alpinus priscus Thenius 1954 was the first member of genus Cuon to be identified in Europe during the Middle Pleistocene. This was followed by Cuon alpinus fossilis Nehring 1890, then Cuon alpinus europaeus Bourguignat 1868 during the Middle and Late Pleistocene. The descent is thought to be C. a. priscus→C. a. fossilis→C. a. europaeus. In comparison, Cuon alpinus antiquus Colbert & Hooijer 1953 was active in the Middle Pleistocene of China, and Cuon alpinus caucasicus Baryshnikov 1996 was active in the Late Pleistocene of the Caucasus. [2]
The European dhole became extinct in much of Europe during the late Würm period, [1] though it may have survived in the Iberian Peninsula until the early Holocene. [3]
Between 650–450 thousand years ago in Europe, the open lands were dominated by Lycaon lycaonoides , while Cuon alpinus priscus preferred forests, highlands, and mountains. The early small wolf Canis mosbachensis coexisted in all of these environments. Between 480–430 thousand years ago the number and range of L. lycaonoides began to fall, and it became extinct across Eurasia between 450– 400 thousand years ago. Between 400–300 thousand years ago, the dhole and the wolf were still similar in size, but the wolf was slowly getting bigger. Between 300–250 thousand years ago, the wolf took over the dominant niche which had once been occupied by L. lycaonoides. Due to competition with the wolf, C. alpinus then decreased in body size and adapted to hunting and living in forests, highlands, and mountains. [4]
The dhole is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. It is genetically close to species within the genus Canis, but distinct in several anatomical aspects: its skull is convex rather than concave in profile, it lacks a third lower molar and the upper molars sport only a single cusp as opposed to between two and four. During the Pleistocene, the dhole ranged throughout Asia, Europe and North America but became restricted to its historical range 12,000–18,000 years ago.
Jackals are medium-sized canids native to Africa and Eurasia. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed jackal and side-striped jackal of sub-Saharan-Africa, and the golden jackal of south-central Europe and Asia. The African golden wolf was also formerly considered as a jackal.
Canis is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and dentition, long legs, and comparatively short ears and tails.
The steppe bison or steppe wisent is an extinct species of bison that was once found on the mammoth steppe where its range included British Isles, Europe, Central Asia, Northern to Northeastern Asia including the Japanese archipelago, Beringia, and central North America, from northwest Canada to Mexico during the Quaternary. This wide distribution is sometimes called the Pleistocene bison belt, compared to the Great bison belt. Three chronological subspecies, Bison priscus priscus, Bison priscus mediator, and Bison priscus gigas, have been suggested.
Speothos is a genus of canid found in Central and South America. The genus includes the living bush dog, Speothos venaticus, and an extinct Pleistocene species, Speothos pacivorus. Unusually, the fossil species was identified and named before the extant species was discovered, with the result that the type species of Speothos is S. pacivorus. S. pacivorus had a larger overall body size and a double-rooted second lower molar. It has been proposed that Speothos originated in the Brazilian highlands sometime during the Pleistocene.
Panthera fossilis, is an extinct species of cat belonging to the genus Panthera, known from remains found in Eurasia spanning the Middle Pleistocene and possibly into the Early Pleistocene. P. fossilis has sometimes been referred to by the common names steppe lion or cave lion, though these names are conventionally restricted to the later related species P. spelaea, to which P. fossilis is probably ancestral.
The Sardinian dhole is an extinct insular canid which was endemic to what is now the Mediterranean islands of Sardinia and Corsica during the Middle-Late Pleistocene. It went extinct at the end of the Pleistocene around the time of human settlement of the islands.
Asoriculus is an extinct genus of terrestrial shrews in the subfamily Soricinae and tribe Nectogalini, native to Europe and North Africa. The best known species, Asoriculus gibberodon, was widespread in Europe from the Late Miocene to the Early Pleistocene. The youngest records of the species date to the end of the Early Pleistocene approximately 846,000 ± 57,000 years ago in the Iberian Peninsula. Another larger species, A. thenii, is sometimes also recognised in the Early Pleistocene of Europe. The species Asoriculus maghrebiensis is known from the Pliocene-Pleistocene boundary of Morocco in North Africa, making it the only known member of Soricinae to have been native to the African continent. Insular species are known from the Mediterranean islands of Sicily, and Corsica-Sardinia including A. corsicanus and A. similis. A. similis likely survived into the Holocene, when it became extinct sometime after human settlement of the islands, with remains apparently being found in Mesolithic and Neolithic aged archaeological sites.
Xenocyon is an extinct group of canids, either considered a distinct genus or a subgenus of Canis. The group includes Canis (Xenocyon) africanus, Canis (Xenocyon) antonii and Canis (Xenocyon) falconeri that gave rise to Canis (Xenocyon) lycanoides. The hypercarnivorous Xenocyon is thought to be closely related and possibly ancestral to modern dhole and the African wild dog, as well as the insular Sardinian dhole.
The latter half of the Late Pleistocene to the beginning of the Holocene saw extinctions of numerous predominantly megafaunal species, which resulted in a collapse in faunal density and diversity across the globe. The extinctions during the Late Pleistocene are differentiated from previous extinctions by the widespread absence of ecological succession to replace these extinct megafaunal species, and the regime shift of previously established faunal relationships and habitats as a consequence. The timing and severity of the extinctions varied by region and are thought to have been driven by varying combinations of human and climatic factors. Human impact on megafauna populations is thought to have been driven by hunting ("overkill") as well as possibly environmental alteration. The relative importance of human vs climatic factors in the extinctions has been the subject of long-running controversy.
Lycaon is a genus of canid which includes the African wild dog and the extinct Lycaon sekowei.
Hypnomys, otherwise known as Balearic giant dormice, is an extinct genus of dormouse (Gliridae) in the subfamily Leithiinae. Its species are considered examples of insular gigantism. They were endemic to the Balearic Islands in the western Mediterranean from the Early Pliocene until their extinction around 4,000 years ago. They first appeared in the fossil record on Mallorca during the Early Pliocene, presumably due to the Messinian salinity crisis causing a connection with mainland Europe. They later spread to Menorca, and a possible molar is also known from Ibiza. Hypnomys became extinct during the Holocene after human arrival on the Balearics. They were one of only three native land mammals to the islands at the time of human arrival, alongside the shrew Nesiotites and goat-antelope Myotragus.
The Petralona Cave also Cave of the Red Stones, a karst formation, is located at 300 m (984 ft) above sea-level on the western foot of Mount Katsika, about 1 km (0.62 mi) east of the village of Petralona, about 35 km (22 mi) south-east of Thessaloniki city on the Chalkidiki peninsula, Greece. The site came to public attention when in 1960 a fossilized archaic human skull was found. The cave had been discovered accidentally only a year earlier (1959) after erosion had left clefts in the rock. "Bejeweled" with impressive stalactite and stalagmite formations and holding an abundance of fossils, the cave soon attracted geologists and paleontologists. After decades of excavations the cave is open to the public and scientific work is documented and presented in an adjacent archaeological museum.
The Ussuri dhole, also known as the Eastern Asiatic dhole and the Chinese dhole, is the nominate subspecies of the dhole wild dog native to Asia. The Ussuri dhole subspecies is originally native to China and sections of Manchuria, the Amur River, the Korean Peninsula and Mongolia; however, it is presumed regionally extinct or extirpated in most of its historical range in the country, and likely found in fragmented populations in the Russian Far East.
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The Noisetier Cave, owing its popular name to the Hazel trees that grow in front of its entrance, is located in a mountainside 145 m (476 ft) atop the Vallée d'Aure in the Ardengost commune, Hautes-Pyrénées department in the region Occitania, Southern France. During systematic excavations since 1992 Middle Paleolithic stone tools and artifacts attributed to the Neanderthal Mousterian culture were discovered among numerous faunal remains.
Mececyon trinilensis, the Trinil dog, is an extinct canid species that lived on the island of Java in Indonesia during the Pleistocene.
Canis mosbachensis, sometimes known as the Mosbach wolf, is an extinct small wolf that once inhabited Eurasia from the Middle Pleistocene era to the Late Pleistocene. It is widely accepted as the ancestor of Canis lupus, the grey wolf.
The steppe brown bear is a disputed extinct subspecies of brown bear that lived in Eurasia during either the Pleistocene or the early Holocene epochs, but its geological age is uncertain. Fossils of the bear have been found in various caves in Slovakia, particularly those of Vazec, Vyvieranie, Lisková, Kupcovie Izbicka, and Okno. It is argued that the subspecies should be rendered invalid, as its geological age is unclear and "its skull is identical to modern U. arctos."
Bison schoetensacki, commonly as the Pleistocene woodland bison or Pleistocene wood bison, was a species of bison that lived until from the Early Pleistocene to at least the early Middle Pleistocene from western Europe to southern Siberia. Its presence in the Late Pleistocene is debated.