Canis lepophagus Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Family: | Canidae |
Genus: | Canis |
Species: | †C. lepophagus |
Binomial name | |
†Canis lepophagus Johnston (1938) [1] | |
Range of Canis lepophagus based on fossil distribution |
Canis lepophagus (Latin: canis: 'dog'; lepus: 'hare' or 'rabbit'; suffix -phagus: '-eating'; hence hare-eating dog) is an extinct species of canid which was endemic to much of North America during the Early Pliocene. [2] It is notable because its lineage is proposed to have led to both wolves and coyotes.
Kurten proposed that the Blancan C. lepophagus derived from smaller Miocene Canis species in North America. It then became widespread across Eurasia where it was either identical to, or closely related with, C. arnensis of Europe. [3] : p241 [4] [5] Wang proposed a linear progression from Eucyon davisi to C. lepophagus to the coyote. [2] : p58
Johnston describes C. lepophagus as having a more slender skull and skeleton than in the modern coyote. [1] : 385 Nowak found that the early populations had small, delicate and narrowly proportioned skulls that resemble small coyotes and appear to be ancestral to C. latrans. [3] : p241 Johnson noted that some specimens found in Cita Canyon, Texas had larger, broader skulls, [1] and along with other fragments Nowak suggested that these were evolving into wolves. [3] : p241 [6]
Tedford disagreed with previous authors and found that its cranio-dental morphology lacked some characteristics that are shared by C. lupus and C. latrans, and therefore there was not a close relationship but it did suggest C. lepophagus was the ancestor of both wolves and coyotes. [7] : p119 Additionally, C. edwardii , C. latrans and C. aureus form together a small clade and because C. edwardii appeared earliest spanning the mid-Blancan (late Pliocene) to the close of the Irvingtonian (late Pleistocene) C. edwardii is proposed as the descendant of C. lepophagus and the ancestor of the coyote and the golden jackal. [2] : p58 [7] : p175, 180
The first fossil record was found in Cita Canyon, Texas. Subsequent discoveries of specimens were found in four other Texas sites, Tonuco Mountain, New Mexico, western Washington [8] Santa Fe River, Florida, [9] Black Ranch in northern California, sites in Nebraska, Idaho, Utah, and Oklahoma.
The coyote is a species of canine native to North America. It is smaller than its close relative, the gray wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecological niche as the golden jackal does in Eurasia. The coyote is larger and was once referred to as the American jackal by a behavioral ecologist. Other historical names for the species include the prairie wolf and the brush wolf.
Canidae is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid. The family includes three subfamilies: the Caninae, the extinct Borophaginae and Hesperocyoninae. The Caninae are known as canines, and include domestic dogs, wolves, coyotes, foxes, jackals and other species.
The dire wolf is an extinct canine. The dire wolf lived in the Americas during the Late Pleistocene and Early Holocene epochs. The species was named in 1858, four years after the first specimen had been found. Two subspecies are recognized: Aenocyon dirus guildayi and Aenocyon dirus dirus. The largest collection of its fossils has been obtained from the Rancho La Brea Tar Pits in Los Angeles.
Canis is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and dentition, long legs, and comparatively short ears and tails.
Chasmaporthetes, also known as hunting or running hyena, is an extinct genus of hyenas distributed in Eurasia, North America, and Africa during the Pliocene-Pleistocene epochs, living from 4.9 million to 780,000 years ago, existing for about 4.12 million years. The genus probably arose from Eurasian Miocene hyenas such as Thalassictis or Lycyaena, with C. borissiaki being the oldest known representative. The species C. ossifragus was the only hyena to cross the Bering land bridge into the Americas, and ranged over what is now Arizona and Mexico during Blancan and early Irvingtonian Land Mammal ages, between 5.0 and 1.5 million years ago.
Borophagus is an extinct genus of the subfamily Borophaginae, a group of canids endemic to North America from the Middle Miocene epoch through the Early Pleistocene epoch 12—1.8 Mya.
Armbruster's wolf is an extinct species that was endemic to North America and lived during the Irvingtonian stage of the Pleistocene epoch, spanning from 1.9 Mya—250,000 years BP. It is notable because it is proposed as the ancestor of one of the most famous prehistoric carnivores in North America, the dire wolf, which replaced it.
Xenocyon is an extinct group of canids, either considered a distinct genus or a subgenus of Canis. The group includes Canis (Xenocyon) africanus, Canis (Xenocyon) antonii and Canis (Xenocyon) falconeri that gave rise to Canis (Xenocyon) lycanoides. The hypercarnivorous Xenocyon is thought to be closely related and possibly ancestral to modern dhole and the African wild dog, as well as the insular Sardinian dhole.
Mount Blanco is a small white hill — an erosional remnant — located on the eastern border of the Llano Estacado within Blanco Canyon in Crosby County, Texas. With Blanco Canyon, it is the type locality of the early Pleistocene Blanco Formation of Texas and Kansas, as well as the Blancan fauna, which occurs throughout North America. Mount Blanco is a Late Blancan age site, and is associated with other Late Blancan sites from Texas such as Red Light and Hudspeth local faunas from Hudspeth County, and the Cita Canyon fauna from Randall County.
Canis edwardii, also known as Edward's wolf, is an extinct species of wolf in the genus Canis which was endemic to North America three million years ago from the Late Blancan stage of the Pliocene epoch and was extinct by the end of the Irvingtonian stage of the Pleistocene epoch.
Canis cedazoensis is an extinct species of smaller canid which was endemic to North America during the Pleistocene epoch, 1.8 Ma—300,000 years ago.
Borophagus dudleyi is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the late Hemphillian of the Miocene epoch through the Pliocene epoch. Borophagus dudleyi existed for approximately 5.4 million years.
Borophagus hilli is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the Late Miocene until the Pliocene.
Borophagus diversidens is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the late Miocene epoch through the Pliocene epoch 4.9—1.8 Ma.
Borophagus secundus is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the Early Miocene epoch through the Late Miocene epoch. Borophagus secundus existed for approximately 17.7 million years.
Eucyon is an extinct genus of medium omnivorous coyote-like canid that first appeared in the Western United States during the late Middle Miocene 10 million years ago. It was the size of a jackal and weighed around 15kg. Its species E. zhoui was one of a number of North American mammals which invaded East Asia around 5–6 million years ago, followed by the genus going extinct 3 million years ago. This genus is proposed to have given rise to genus Canis 6 million years ago.
Canis nehringi is an extinct species of canid. Canis gezi, a poorly known small wolf from the Ensenadan of South America, appears to have given rise to Canis nehringi, a Lujanian species from Argentina. The species was first described by Florentino Ameghino in 1902.
The evolution of the wolf occurred over a geologic time scale of at least 300 thousand years. The grey wolf Canis lupus is a highly adaptable species that is able to exist in a range of environments and which possesses a wide distribution across the Holarctic. Studies of modern grey wolves have identified distinct sub-populations that live in close proximity to each other. This variation in sub-populations is closely linked to differences in habitat – precipitation, temperature, vegetation, and prey specialization – which affect cranio-dental plasticity.
Canis arnensis, the Arno River dog, is an extinct species of canine that was endemic to Mediterranean Europe during the Early Pleistocene. The Arno River dog has been described as a small jackal-like dog. Its anatomy and morphology relate it more to the modern golden jackal than to the larger Etruscan wolf of that time. It is probably the ancestor of modern jackals.
Canis mosbachensis is an extinct wolf that once inhabited Europe 600,000—420,000 years ago. The Mosbach wolf was a short-legged carcass feeder adapted for scavenging megafauna on the mammoth steppe. The Mosbach wolf is proposed as the ancestor of the grey wolf Canis lupus but some mammalogists have assigned it as the subspecies Canis lupus mosbachensis.