Borophagus littoralis [1] | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Family: | Canidae |
Genus: | † Borophagus |
Species: | †B. littoralis |
Binomial name | |
†Borophagus littoralis VanderHoof, 1931 | |
Borophagus littoralis ("coastal glutton") is an extinct species of the genus Borophagus of the subfamily Borophaginae , a group of canids endemic to North America from the early Miocene epoch (23.3 Mya) through the Pliocene epoch (4.9 Mya). Borophagus littoralis existed for approximately 18.4 million years. [2]
Borophagus littoralis was named by Cope in 1892 and is considered synonymous with Osteoborus diabloensis. Borophagus littoralis, like other Borophaginae, are loosely known as "bone-crushing" or "hyena-like" dogs. Though not the most massive borophagine by size or weight, it had a more highly evolved capacity to crunch bone than earlier, larger genera such as Epicyon , which seems to be an evolutionary trend of the group (Turner, 2004). During the Pliocene epoch, Borophagus began being displaced by Canis genera such as Canis edwardii and later by Canis dirus . Early species of Borophagus were placed in the genus Osteoborus until recently, but the genera are now considered synonyms. [1] Borophagus littoralis possibly led a hyena-like lifestyle scavenging carcasses of recently dead animals.
Typical features of this genus are a bulging forehead and powerful jaws; it was probably a scavenger. [3] Its crushing premolar teeth and strong jaw muscles would have been used to crack open bone, much like the hyena of the Old World. The adult animal is estimated to have been about 80 cm in length, similar to a coyote, although it was much more powerfully built. [4]
Canidae is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid. The family includes three subfamilies: the Caninae, and the extinct Borophaginae and Hesperocyoninae. The Caninae are known as canines, and include domestic dogs, wolves, coyotes, foxes, jackals and other species.
Borophagus is an extinct genus of the subfamily Borophaginae, a group of canids endemic to North America from the Middle Miocene epoch through the Early Pleistocene epoch 12—1.8 Mya.
Aelurodon is an extinct canid genus of the subfamily Borophaginae which lived from the Barstovian land mammal age of the middle Miocene to the late Miocene epoch. Aelurodon existed for approximately 10.7 million years.
The extinct Borophaginae form one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Hesperocyoninae and extant Caninae. Borophaginae, called "bone-crushing dogs", were endemic to North America during the Oligocene to Pliocene and lived roughly 34—2.5 million years ago, existing for about 33.5 million years.
Epicyon is a large, extinct, canid genus of the subfamily Borophaginae, native to North America. Epicyon existed for about 15 million years from the Hemingfordian age of the Early Miocene, to the Hemphillian of the Late Miocene. Epicyon haydeni is the largest known canid of all time, with the type species reaching 2.4 m (7.9 ft) in length, 90 cm (35 in) in shoulder height and approximately 100–125 kg (220–276 lb) in body mass. The largest known humerus specimen belonged to an individual weighing up to 170 kg (370 lb).
Hesperocyon is an extinct genus of canids that was endemic to North America, ranging from southern Canada to Colorado. It appeared during the Uintan age, –Bridgerian age (NALMA) of the Mid-Eocene– 42.5 Ma to 31.0 Ma. (AEO). Hesperocyon existed for approximately 11.5 million years.
Cynodesmus is an extinct genus of omnivorous canine which inhabited North America during the Oligocene living from 33.3—-26.3 Ma and existed for approximately 7 million years.
Caninae is the only living subfamily within Canidae, alongside the extinct Borophaginae and Hesperocyoninae. They first appeared in North America, during the Oligocene around 35 million years ago, subsequently spreading to Asia and elsewhere in the Old World at the end of the Miocene, some 7 million to 8 million years ago.
Tomarctus is a canid genus of the extinct subfamily Borophaginae which inhabited most of North America during the late Early Miocene to the Early Barstovian age of the Middle Miocene. Tomarctus existed for approximately 6.83 million years.
Cynarctus is an extinct genus of the Borophaginae subfamily of canids native to North America. The genus was first founded by W. D. Matthew in 1901, based from a pair of lower jaws, Cynarctus saxitilis, found in the Pawnee Creek Beds of Colorado. It lived during the Middle to Late Miocene 16.0—10.3 mya, existing for approximately 5.7 million years. Fossils have been uncovered in Colorado, California, Maryland, western Nebraska, and Texas. It was likely an omnivore, and lacked the bone-cracking adaptations found in some later borophagines. Newer findings have proved the genus to be described as a large dog-like raccoon, a result from combining characteristics from Canidae with Procyonidae.
Borophagini is a clade or tribe of the subfamily Borophaginae. This is an extinct group of terrestrial canids that were endemic and widespread throughout North America and Central America which lived during the Geringian stage of the Oligocene epoch to the Zanclean age of the Early Pliocene living 30.8—3.6 Mya existing approximately 27.2 million years.
Borophagina is a subtribe of the Borophaginae, a group of extinct canids. They inhabited much of North America from the Early Miocene to the Zanclean stage of the Pliocene, 20.6—3.6 Mya, and existed for approximately 17 million years.
Canis edwardii, also known as Edward's wolf, is an extinct species of wolf in the genus Canis which was endemic to North America three million years ago from the Late Blancan stage of the Pliocene epoch and was extinct by the end of the Irvingtonian stage of the Pleistocene epoch.
Borophagus parvus is an extinct species of the genus Borophagus, of the subfamily Borophaginae, a group of canids endemic to North America from the late Hemphillian of the Miocene epoch through the Pliocene epoch 10.3—4.9 Ma, existing for approximately 5.4 million years.
Borophagus dudleyi is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the late Hemphillian of the Miocene epoch through the Pliocene epoch. Borophagus dudleyi existed for approximately 5.4 million years.
Borophagus hilli is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the Late Miocene until the Pliocene.
Borophagus diversidens is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the late Miocene epoch through the Pliocene epoch 4.9—1.8 Ma.
Borophagus orc is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the 10.3 Mya to 4.9 Mya. Borophagus orc existed for approximately 5.4 million years.
Borophagus pugnator is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the early Miocene epoch through the late Miocene epoch 23.3—5.3 Ma. Borophagus pugnator existed for approximately 18 million years.
Borophagus secundus is an extinct species of the genus Borophagus of the subfamily Borophaginae, a group of canids endemic to North America from the Early Miocene epoch through the Late Miocene epoch. Borophagus secundus existed for approximately 17.7 million years.