Formica rufibarbis | |
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Formica rufibarbis worker | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hymenoptera |
Family: | Formicidae |
Subfamily: | Formicinae |
Genus: | Formica |
Species: | F. rufibarbis |
Binomial name | |
Formica rufibarbis Fabricius, 1793 | |
Formica rufibarbis is a European formicine ant of the Formica fusca group. In the classification by Auguste Forel, it is treated in the subgenus Serviformica . F. rufibarbis is subject to a Species Action Plan (SAP) in England, where it is known from only two locations, although it is not considered to be at risk on continental Europe.[ citation needed ]
The name Formica rufibarbis was first given to this ant by Lord Avebury in Britain in his 1881 work Ants, Bees and Wasps although the species had been earlier misidentified as F. cunicularia by Frederick Smith in 1851.[ citation needed ]
The Red-barbed Ant is readily identified by its relatively large size and distinctive coloration of a blackish head and thorax, contrasting with a light reddish thorax. Small dark workers do occur and may be mistaken for F. fusca , although there is always a degree of colouration between thorax and abdomen. [1] Workers can also be confused with F. cunicularia which does not have hairs on the thorax. [2]
Colonies usually contain one to three queens, although the occurrence of gynaecoid or egg laying workers has also been recorded. Queens found colonies in the same manner as other ants from the Formica fusca group. Eggs are first laid early in the new year, and colonies reach a maximum size of around 500 workers. Alates emerge in late June to early July.[ citation needed ]
It is locally common throughout continental Europe, and ranges from Portugal to Western Siberia. it nests in short, lowland grass and heather or maritime heath overlying loose or sandy soils. [3]
In Britain, the species has always been scarce, confined to heaths in Surrey and the Isles of Scilly, where it is sometimes known as the "St Martin's Ant". In the 1927 edition of British Ants: their life histories and classification, Donisthorpe gives its distribution as being confined to Ripley, Chobham, Reigate and Weybridge. In 2004 there were only four nests in Surrey. It was once found in Cornwall at Whitsand Bay but has not been recorded since 1907. In the Isles of Scilly it is found on the islands of Great Ganilly, Nornour, St Martin's and Teän. [2] As of 2015, F. rufibarbis is now confined to one known colony on mainland Britain, nesting on the edge of a Heath, the location is kept a closely guarded secret due to the risk posed by curious members of the public. However the nest is also at risk from F. sanguinea, a slavemaker ant, which currently nests only a few hundred meters away.
F. rufibarbis nests completely within the ground, usually in sandy banks, and nest chambers situated about a foot beneath the surface are accessible only from a single entrance. This makes the locating of colonies very difficult, so it is possible that the small numbers of recorded colonies constitute an under-representation. Workers forage singly and Donisthorpe observed: "The workers are very audacious and will even endeavour to rob F. rufa of its prey – holding on and pulling – and the moment the rufa lets go, to get a better grip, or to attack the rufibarbis, the latter swiftly decamps with the prize." Workers also possess a remarkable sense of sight, and will proceed to their nest entrance in a dead straight line even if major obstacles are placed to disrupt their path. Donisthorpe describes this phenomenon thus: "On July 12, 1913, having observed several rufibarbis workers running about on a path near a sandy bank at Weybridge, I endeavoured to find their nest, and commenced to pull up handfuls of herbage on the top of the bank, which I let fall on the slope. I then saw a worker approaching with a fly in its jaws and start to mount the bank, and as the scattered herbage was directly in its way, I feared the ant might be diverted from its nest, but when it reached the obstacle it never hesitated for a moment, but running straight over it in a direct line, entered its nest on the top of the bank, which I was thus enabled to find."[ citation needed ]
Like other Serviformica, this species is subject to raids by dulotic species such as Formica sanguinea and Polyergus rufescens where their ranges coalesce. In Britain this only takes place at Chobham in the case of the former (the latter does not occur in the country).[ citation needed ]
Formica exsecta is a species of ant found from Western Europe to Asia.
The erratic ant is a species of dolichoderine ant first described in 1789 by Latreille.
Horace St. John Kelly Donisthorpe was an eccentric British myrmecologist and coleopterist, memorable in part for his enthusiastic championing of the renaming of the genus Lasius after him as Donisthorpea, and for his many claims of discovering new species of beetles and ants.
Formica is a genus of ants of the family Formicidae, commonly known as wood ants, mound ants, thatching ants, and field ants. Formica is the type genus of the Formicidae, and of the subfamily Formicinae. The type species of genus Formica is the European red wood ant Formica rufa. Ants of this genus tend to be between 4 and 8 mm long. Ants belonging to the Formica genus possess a single knob or bump located between their thorax and abdomen. These ants primarily feed on honeydew, a sugary liquid produced by aphids. Interestingly, Formica ants appear to take on a shepherding role with smaller aphids, relocating them to different parts of plants to ensure a continuous food source for the aphids. By doing so, the ants can establish a relatively sustainable honeydew supply for both themselves and their colony.
The Formica rufa group is a subgeneric group within the genus Formica, first proposed by William Morton Wheeler. This group contains the mound-building species of Formica commonly termed "wood ants" or "thatch-mound ants", which build prominent nests consisting of a mound of grass, litter, or conifer needles. The species Formica rufa or the red wood ant is the type species of this subgroup.
Formica rufa, also known as the red wood ant, southern wood ant, or horse ant, is a boreal member of the Formica rufa group of ants, and is the type species for that group, being described already by Linneaus. It is native to Eurasia, with a recorded distribution stretching from the middle of Scandinavia to the northern Iberia and Anatolia, and from Great Britain to Lake Baikal, with unconfirmed reportings of it also to the Russian Far East. There are claims that it can be found in North America, but this is not confirmed in specialised literature, and no recent publication where North American wood ants are listed mentions it as present, while records from North America are all listed as dubious or unconfirmed in a record compilation. Workers head and thorax are colored red and the abdomen brownish-black, usually with a dorsal dark patches on the head and promensonotum, although some individuals may be more uniform reddish and even have some red on the part of the gastern facing the body. In order to separate them from closely related species, specimens needs to be inspected under magnification, where difference in hairyness are among the telling characteristics, with Formica rufa being more hairy than per example Formica polyctena but less hairy than Formica lugubris. Workers are polymorphic, measuring 4.5–9 mm in length. They have large mandibles, and like many other ant species, they are able to spray formic acid from their abdomens as a defence. Formic acid was first extracted in 1671 by the English naturalist John Ray by distilling a large number of crushed ants of this species. These ants primarily eat honeydew from aphids. Some groups form large networks of connected nests with multiple queen colonies, while others have single-queen colonies.
Formica cunicularia is a species of ant found all over Europe. They are especially common in western Europe and southern England, but they can be found from southern Scandinavia to northern Africa and from Portugal to the Urals. In England, Donisthorpe records the species as having occurred as far north as Bewdley in Worcestershire. In Formica cunicularia, the worker is an ashy grey black color and is usually 4.0–6.5 mm long. The males are found to have a uniformly dark body and are 8.0–9.0 mm long. The queen is yellowish red to dark black and is 7.5–9.0 mm.
Polyergus is a small genus of ants with 14 described species. They are also referred to by the names "slave-raiding ants" and "Amazon ants". They are characterized by their habit of raiding nests for workers.
Formica fusca is a black-colored ant commonly found throughout Europe as well as parts of Southern Asia and Africa. It has the common names silky ant or dusky ant. The range within the palaearctic region extends from Portugal in the west to Japan in the east and from Italy in the south to Fennoscandia in the north. Populations from North America have been split off as a separate species, Formica subaenescens. F. fusca nests are usually found in rotten tree stumps or under stones in clearcut areas and along woodland borders and hedgerows.
Formica lugubris, also known as the hairy wood ant is commonly found in wooded upland areas across northern Eurasia. Colonies construct large thatched mound nests occupied by thousands of workers, and one or more queens. Workers look similar to other species of wood ants, but Formica lugubris workers can be identified by a fringe of hairs that reaches down to their eyes and prominent hairs between the facets of their compound eyes. Workers can reach sizes of up to 9 mm long; queens are larger, reaching 12 mm long.
Formica pratensis, the black-backed meadow ant, is a species of European red wood ant in the family Formicidae.
Teän is an uninhabited island to the north of the Isles of Scilly archipelago between Tresco, 1.5 kilometres (0.9 mi) to the west, and St Martin's, 300 metres (330 yd) to the east. Approximately 16 hectares in area, the island consists of a series of granite tors with the highest point, Great Hill, rising to 40 metres (130 ft) at its eastern end. The low-lying land is overlain with glacial till and outwash gravels with glacial erratics abundant on the north coast beaches, which indicates the southern limit of outwash from an ice sheet for which it is designated a Geological Conservation Review site.
Formica sanguinea, or blood-red ant, is a species of facultative slave-maker ant in the genus Formica characterized by the ability to secrete formic acid. It ranges from Central and Northern Europe through Russia to Japan, China, the Korean Peninsula, Africa and also the United States. This species is coloured red and black with workers up to 7 mm long.
Formica cinerea is a species of ant in the family Formicidae.
Formica pacifica is an ant species occupying the Pacific Coast of the United States and British Columbia, Canada. This species is a member of the Formica fusca species group. Workers are quite fast and aggressive. This species engages in wars with neighboring colonies. F. pacifica is present in both forests and human settlements. Colonies of this species can grow up to a few thousand workers.
Slave-making ants or slaver ants are brood parasites that capture broods of other ant species to increase the worker force of their colony. After emerging in the slave-maker nest, slave workers work as if they were in their own colony, while parasite workers only concentrate on replenishing the labor force from neighboring host nests, a process called slave raiding.
Polyergus rufescens is a species of slave-making ant native to southern Europe and parts of Asia, commonly referred to as the European Amazon ant or as the slave-making ant. It is an obligatory social parasite, unable to feed itself or look after the colony and reliant on ants of another species to undertake these tasks. To replenish these servant ants, it raids nearby ant colonies and carries home pupae and larvae, and these are reared to provide future workers for the colony. A newly mated female P. rufescens needs to make its way into one of these "host" nests, kill the host queen, and be accepted by the host workers in her place.
Formica incerta is a species of ant found in eastern North America. It is the most common species of Formica in many areas, and excavates underground nests with small entrance holes. Its diet includes nectar produced by extrafloral nectaries and honeydew, which it obtains from aphids and treehoppers. It is the main host for the slave-making ant Polyergus lucidus. F. incerta was first described by Italian entomologist Carlo Emery in 1893. Its specific name comes from the Latin incertus meaning "uncertain" and seems particularly apt given the subsequent uncertainty as to the validity of the species and the difficulty in distinguishing this ant from other species living in the same area.
The Isles of Scilly are an archipelago 45 km (28 mi) off Land's End, Cornwall. Little of the fauna on, above or in the seas surrounding the isles was described prior to the 19th century, when birds and fish started to be described. Most records of other animals date from the 20th century onwards.