Galax | |
---|---|
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Clade: | Asterids |
Order: | Ericales |
Family: | Diapensiaceae |
Genus: | Galax Sims |
Species: | G. urceolata |
Binomial name | |
Galax urceolata | |
Galax, the wandplant, wandflower, or beetleweed, is a genus in the flowering plant family Diapensiaceae, containing a single species, Galax urceolata (syn. G. rotundifolia, G. aphylla). It is native to the southeastern United States from Massachusetts and New York south to northern Alabama, growing mainly in the Appalachian Mountains at altitudes of up to 1,500 m, where it grows in shaded places in forests. Galax urceolata can occur at multiple ploidy levels, an individual may be a diploid (2x), triploid (3x), or autotetraploid (4x) (autopolyploid). [1] The cytotypes are neither morphologically nor geographically distinct, though there are slight climatic differences between the diploid and autotetraploid types. [2] Outcrossing is likely to occur among cytotypes as well. [3]
It is an evergreen herbaceous perennial plant growing to 30–45 cm (rarely 75 cm) tall, with a rosette of leathery leaves, which turn brown during winter. The leaves are a rounded cardioid (heart) shape, 2.5–7.5 cm diameter, rarely up to 15 cm, with a serrated margin with rounded "teeth". The flowers are produced in late spring to early summer, white in color and on a single spike-like raceme 15–25 cm long on top of a 20–50 cm tall stem. Each individual flower has five petals, and is up to 4 mm (5⁄32 in) in diameter. The fruit is a small capsule containing numerous seeds.
The genus name Galax comes from the Greek word "gala" which means "milk", [4] describing Galax's white flowers.
Around 1730, Galax was collected by John Clayton, due to his friendship with Mark Catesby, an English naturalist who had just arrived in Virginia. Based on Catesby's recommendation, Clayton ended up sending his specimens to Jan Frederik Gronovius, a Dutch botanist. [5] In 1739, Gronovius published The Flora of Virginia, where "Anonymos or Belvedere" is the plant long known as Galax aphylla. Clayton provided Gronovius with four samples, all which were destroyed in a series of unfortunate events. [5]
Many years later, John Mitchell assumed he had collected Galax aphylla and took a ship to bring the specimen to Carl Linnaeus; however, his boat was attacked by pirates, who took all the specimens as part of their plunder. Prior to the voyage, he had sent descriptions of all the specimens to colleagues in Europe. When he arrived there, he was able to procure one of his descriptions for Linnaeus. Despite Linnaeus never seeing this specimen of “Galax”, he agreed with Mitchell's description; however, the specimen described was, in fact, " Nymophila ", thus invalidating the name Galax aphylla with respect to the International Code of Nomenclature of Plants.
Between March 1803 and September 1804, Galax was rediscovered and renamed five times. Any time the new name referred to Mitchell's description, it was invalidated. The name that is considered valid, since it pre-dates all other names and follows the rules of nomenclature, was Pyrola urceolata Poir., named by Jean Louis Marie Poiret. Despite the validity of this name, Pyrola never caught on. [5] Around this time Galax urceolata was cultivated in Europe; many descriptions came from the cultivated line (presumably sent by John Clayton), and the plant referred to as "Galax" was gaining popularity. In 1972, Brummitt argued that the genus name should remain Galax, but the specific epithet should be urceolata; therefore, he renamed it Galax urceolata (Poir.) Brummitt. [5] However, many herbaria still label this specimen with its original invalid name, Galax aphylla.
The type specimen was collected by André Michaux, who had his horses stolen on the journey to collect Galax with John Clayton. Since the prior specimens had been destroyed and the name invalidated, this new specimen would be considered the holotype.
Name | Authority | Date | Description |
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Erythrorhiza rotundifolia | André Michaux | March 1803 | Noted as a synonym of Galax aphylla Linn. therefore invalidated. |
Pyrola urceolata | Jean Louis Marie Poiret | January 1804 | Based on a plant in cultivation in France, in the garden of M. Lemonier. |
Blandfordia cordata | Henry C. Andrews | February 1804 | Blandfordia was used for a genus in Liliaceae in 12/1804, and the genus name is still conserved. This name is therefore invalidated. |
Galax aphylla | John Sims | June 1804 | Stated synonymy with Erythrorhiza rotundifolia of Michaux, Blandfordia cordata of Andrews, Belvedere of Clayton & Gronovius, Viticella of Mitchell, and Pyrola from Poiret. - invalidated |
Solenandria cordifolia | E. P. Ventenat | September 1804 | Pointed out that Belvedere of Clayton & Gronovius and Viticella of Mitchell were not the same plant. |
The leaves are often harvested for the floristry industry; concern has been expressed over excessive exploitation, and collection is now restricted in many areas. It has also been used in herbalism to treat cuts and kidney ailments. It is occasionally grown as an ornamental plant in gardens.
The independent city of Galax, Virginia, as well as Galax, North Carolina are named after this plant.
Polyploidy is a condition in which the cells of an organism have more than two paired sets of (homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one from each of two parents; each set contains the same number of chromosomes, and the chromosomes are joined in pairs of homologous chromosomes. However, some organisms are polyploid. Polyploidy is especially common in plants. Most eukaryotes have diploid somatic cells, but produce haploid gametes by meiosis. A monoploid has only one set of chromosomes, and the term is usually only applied to cells or organisms that are normally diploid. Males of bees and other Hymenoptera, for example, are monoploid. Unlike animals, plants and multicellular algae have life cycles with two alternating multicellular generations. The gametophyte generation is haploid, and produces gametes by mitosis; the sporophyte generation is diploid and produces spores by meiosis.
Acorus is a genus of monocot flowering plants. This genus was once placed within the family Araceae (aroids), but more recent classifications place it in its own family Acoraceae and order Acorales, of which it is the sole genus of the oldest surviving line of monocots. Some older studies indicated that it was placed in a lineage, that also includes aroids (Araceae), Tofieldiaceae, and several families of aquatic monocots. However, modern phylogenetic studies demonstrate that Acorus is sister to all other monocots. Common names include calamus and sweet flag.
Jan Frederik Gronovius was a Dutch botanist notable as a patron of Linnaeus.
Mark Catesby was an English naturalist who studied the flora and fauna of the New World. Between 1729 and 1747, Catesby published his Natural History of Carolina, Florida and the Bahama Islands, the first published account of the flora and fauna of North America. It included 220 plates of birds, reptiles, amphibians, fish, insects, mammals and plants.
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Claytonia is a genus of flowering plants native to Asia, North America, and Central America. The vitamin-rich leaves can be eaten raw or cooked, and the tubers can be prepared like potatoes.
Jane Colden was an American botanist, described as the "first botanist of her sex in her country" by Asa Gray in 1843. Although not acknowledged in contemporary botanical publications, she wrote a number of letters resulting in botanist John Ellis writing to Carl Linnaeus of her work applying the Linnaean system of plant identification to American flora, for which botanist Peter Collinson stated "she deserves to be celebrated". Contemporary scholarship maintains that she was the first female botanist working in America, which ignores, among others, Maria Sibylla Merian or Catherine Jérémie. Colden was respected as a botanist by many prominent botanists including John Bartram, Peter Collinson, Alexander Garden, and Carl Linnaeus. Colden is most famous for her untitled manuscript, housed in the British Museum, in which she describes the flora of the Hudson Valley in the Newburgh region of New York state, including ink drawings of 340 different species.
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John Clayton (1694/5–1773) was an Anglican minister in and for decades clerk for Gloucester County in the Colony of Virginia who is today best known as a plant collector and botanist. He may be confused with several distant family members, including Rev. John Clayton who served as minister at Jamestown (1682–1684) and conducted various scientific experiments, before returning to England.
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Iris aphylla is a species in the genus Iris, it is also in the subgenus Iris, and in the section Iris. It is a rhizomatous perennial, from Asia to Europe. It is found in Azerbaijan, Russian Federation, Czech Republic, Germany, Hungary, Poland, Belarus, Ukraine, Bulgaria, Albania, Former Yugoslavia, Italy, Romania and France. It has dark green or bright green, sword-shaped, long grass-like leaves, that die/fade away in the winter. It also has a slender stem, with several branches and green and purplish spathes. It has 3–5 large flowers, in shades of bright purple, purple, violet, dark blue, blue-violet and dark violet, which bloom between spring and early summer. Occasionally, they re-bloom in the autumn, before the seed capsule is formed. It is cultivated as an ornamental plant in temperate regions. There is one known subspecies Iris aphylla subsp. hungarica Hegi.
Iris aphylla subsp. hungarica is a plant subspecies in the genus Iris, it is also in the subgenus Iris. It is a subspecies of Iris aphylla. It is a rhizomatous perennial, from the Carpathian Mountains and Pannonian Basin. Found within the countries of Romania, Hungary, Slovakia, Ukraine, Moldova, and Italy. It has curved, pointed, thin leaves, slender stem, several large flowers in shades of purple, or dark purple, or violet-blue. It is a rare plant, that is protected by environmental law in the various countries of Europe.
Iris benacensis is a plant species in the genus Iris, it is also in the subgenus Iris. It is a rhizomatous perennial, from Italy. It has similar sized leaves and stem, and blue-purple shaded flowers, that have a white, blue and yellow beard. It was once classified as a synonym of Iris aphylla, before being re-classified as a species in its own right, although some sources still call it a synonym. It is cultivated as an ornamental plant in temperate regions.
Iris perrieri is a plant species in the genus Iris; it is also in the subgenus Iris. It is a rhizomatous perennial, from the Savoy Alps in southern France and recently in Italy. It has green, deeply ribbed, sickle shaped leaves, a slender stem with a branch, 1–3 scented flowers that are violet or purple, with a white or pale blue beard. It is rarely cultivated as an ornamental plant in temperate regions, due to its rarity in the wild. It was once thought to be a form of Iris aphylla, before cell (chromosome) analysis determined it to be a separate species.