Hackelia

Hackelia
Hackelia diffusa
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Boraginales
Family:	Boraginaceae
Subfamily:	Boraginoideae
Genus:	Hackelia Opiz (1839)
Type species
Hackelia deflexa Opiz
Species
About 54 (see text)
Synonyms [1]
AustrocynoglossumPopov ex R.Mill. (1989)

Hackelia (stickseeds) is a genus of plants in the borage family, Boraginaceae. It includes 54 species found in North America, western South America, temperate Eurasia, and Australia. ^[1] 12 species are native to California. ^[2]

The genus was named after Josef Hackel, a Czech botanist. The common name, stickseed, refers to the tendency of the barbed nutlets to stick to animal fur. ^[3]

Species

54 species are currently accepted: ^[1]

• Hackelia amethystina J.T.Howell - amethyst stickseed – California
• Hackelia andicola (K.Krause) Brand – Peru
• Hackelia bella (J.F.Macbr.) I.M.Johnst. - greater showy stickseed – northwestern California and southwestern Oregon
• Hackelia besseyi (Rydb.) J.L. Gentry - Bessey's stickseed – Arizona, Colorado, New Mexico and Texas
• Hackelia bhutanica R.R.Mill – eastern Nepal to northwestern Arunachal Pradesh
• Hackelia brachytuba (Diels) I.M.Johnst. – Nepal to central China
• Hackelia brevicula (Jeps.) J.L.Gentry - Poison Canyon stickseed – eastern California
• Hackelia californica (A.Gray) I.M.Johnst. - California stickseed – northern California and west-central Oregon
• Hackelia ciliata (Douglas ex Lehm.) I.M.Johnst. - British Columbia, Idaho, and Washington
• Hackelia cinerea (Piper) I.M.Johnst. - gray stickseed – Washington, Idaho, and Montana
• Hackelia cronquistii J.L.Gentry - Cronquist's stickseed – Oregon and Idaho
• Hackelia cusickii (Piper) Brand - Cusick's stickseed – Oregon, northern California, and northwestern Nevada
• Hackelia davisii Cronquist – Idaho
• Hackelia deflexa (Wahlenb.) Opiz – northern Eurasia and northern North America
• Hackelia difformis (Y.S.Lian & J.Q.Wang) Riedl – south-central China and Tibet
• Hackelia diffusa (Douglas ex Lehm.) I.M.Johnst. - Spreading stickseed – British Columbia, Washington, Oregon
• Hackelia floribunda (Lehm.) I.M.Johnst. - manyflower stickseed – western and central Canada and United States
• Hackelia gracilenta I.M.Johnst. – Colorado
• Hackelia heliocarpa (Brand) Brand – Mexico (Chihuahua)
• Hackelia hendersonii (Piper) Brand – Washington, Oregon, and Montana
• Hackelia hintoniorum (B.L.Turner) Sutorý – Mexico (Oaxaca)
• Hackelia hirsuta (Wooton & Standl.) I.M.Johnst. – New Mexico
• Hackelia hispida I.M.Johnst. - showy stickseed – Washington, Oregon, Idaho
• Hackelia ibapensis L.M.Shultz & J.S.Shultz – Utah
• Hackelia latifolia (R.Br.) Dimon & M.A.M.Renner – eastern Australia and Tasmania
• Hackelia leonotis I.M.Johnst. – northeastern and central Mexico
• Hackelia macrophylla (Brand) I.M.Johnst. – northern Pakistan and western Himalaya
• Hackelia meeboldii Brand – western Himalaya
• Hackelia mexicana (Schltdl. & Cham.) I.M.Johnst. – Mexico to Venezuela and Peru
• Hackelia micrantha (Eastw.) J.L.Gentry - Jessica sticktight – western Canada and United States
• Hackelia mundula (Jeps.) Ferris - pink stickseed – southwestern Oregon to central California
• Hackelia murgabica Czukav. – Tajikistan
• Hackelia nervosa (Kellogg) I.M.Johnst. - Sierra stickseed – California and northwestern Nevada
• Hackelia obtusifolia R.R.Mill – Bhutan
• Hackelia ophiobia R.L.Carr – southeastern Oregon, southwestern Idaho, and northern Nevada
• Hackelia parviflora (K.Krause) Brand – Peru (Junín)
• Hackelia patens (Nutt.) I.M.Johnst. - spotted stickseed – west-central United States
• Hackelia pinetorum (Greene ex A.Gray) I.M.Johnst. – Arizona, New Mexico, Texas, and northern Mexico
• Hackelia popovii Czukav. – Tajikistan
• Hackelia rattanii (Brand) Brand – Mexico (Baja California)
• Hackelia revoluta (Ruiz & Pav.) I.M.Johnst. – Colombia to northwestern Argentina
• Hackelia sessilifructa (Y.S.Lian & J.Q.Wang) Ovczinnikova – Xinjiang
• Hackelia setosa (Piper) I.M.Johnst. - bristly stickseed – northern California and southwestern Oregon
• Hackelia sharsmithii I.M.Johnst. - Sharsmith's stickseed – central California and central Nevada
• Hackelia skutchii I.M.Johnst. – Guatemala and Costa Rica
• Hackelia stewartii I.M.Johnst. – northern Pakistan and western Himalaya
• Hackelia stricta I.M.Johnst. – southwestern Mexico to Veracruz
• Hackelia suaveolens (R.Br.) Dimon & M.A.M.Renner – eastern and southern Australia and Tasmania
• Hackelia taylorii Harrod, Malmquist & R.L.Carr – Washington
• Hackelia uncinata (Royle ex Benth.) C.E.C.Fisch. – northeastern Pakistan to Himalaya and southern China
• Hackelia ursina (Greene ex A.Gray) I.M.Johnst. – southeastern Arizona, southwestern New Mexico, and northern Mexico (Sonora)
• Hackelia velutina (Piper) I.M.Johnst. - velvet stickseed – north-central and central California and northwestern Nevada
• Hackelia venusta (Piper) H.St.John - lesser showy stickseed – central Washington
• Hackelia virginiana (L.) I.M.Johnst. - beggar's lice – central and eastern United States and eastern Canada

References

1. Hackelia Opiz. Plants of the World Online . Retrieved 17 June 2023.
2. Calflora Database: Hackelia species index in California. Calflora. Retrieved 17 June 2023.
3. Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 24. ISBN   0-87842-280-3. OCLC   25708726.

External links

• USDA Plants Profile
• Calflora Database: Hackelia species index in California
• Jepson Manual eFlora (TJM2) treatment of Hackelia in California