| Proplatynotia Temporal range: Late Cretaceous, | |
|---|---|
Scientific classification  | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Order: | Squamata |
| Superfamily: | Varanoidea |
| Genus: | † Proplatynotia Borsuk-Bialynicka, 1984 |
| Species | |
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Proplatynotia is an extinct genus of varanoid lizard from the Late Cretaceous of Mongolia. Fossils have been found in the Barun Goyot Formation, which is mid-Campanian in age. The type and only species, P. longirostrata, was named in 1984.
Proplatynotia was named in 1984 from a single holotype skull cataloged as ZPAL MgR-I/68. The skull is mostly complete except for damage in the posterior area. Rounded plates of osteoderms cover the top surface and would have been covered in scales in life.
Compared to other Cretaceous varanoids, Proplatynotia was medium-sized with a skull about 4 centimetres (1.6 in) long. Its skull is long and slender, giving the type species P. longirostrata its name, which means "long snout." Bones in the back of the skull called parietals are enlarged, indicating that it may have had larger jaw muscles and a stronger bite than most monitor lizards. [1] Proplatynotia has many primitive varanoid features, including nostrils that are placed near the tip of the snout. The nostrils have expanded and moved backward in later varanoids, separating the maxilla and nasal bone. In Proplatynotia, these bones still touch. Other characteristic features of Proplatynotia include:
When Proplatynotia was named in 1984, it was classified as a platynotan lizard. Platynota is the group that includes monitor lizards and sometimes snakes and mosasaurs. It includes all members of the superfamily Varanoidea. Along with other Cretaceous lizards from Mongolia like Parviderma and Gobiderma , Proplatynotia was called a necrosaurian. Necrosaurians were an evolutionary grade of varanoid lizards that had generalized morphologies, lacking the derived features of monitor lizards and snakes. They were the most primitive platynotans. [3]
Since the first description of Proplatynotia, some features have been recognized that separate it from other platynotans. The teeth of Proplatynotia are smooth and lack plicidentine, folded layers of dentine around the center pulp cavity of the tooth. Plicidentine, while rare in land-living vertebrates, is characteristic of monitor lizards and their closest extinct relatives. The lack of plicidentine in Proplatynotia led to an early hypothesis that it was not a platynotan, but evolved platynotan features through convergence. [3]
Despite its non-platynotan features, Proplatynotia is usually classified as one of the most basal platynotans. In most recent studies, Platynotia includes monitor lizards, helodermatids, and their close extinct relatives, but not snakes. Below is a cladogram from a 2008 phylogenetic study showing the relationships of Proplatynotia: [4]
| Platynota |
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Garudimimus is a genus of ornithomimosaur that lived in Asia during the Late Cretaceous. The genus is known from a single specimen found in 1981 by a Soviet-Mongolian paleontological expedition in the Bayan Shireh Formation and formally described in the same year by Rinchen Barsbold; the only species is Garudimimus brevipes. Several interpretations about the anatomical traits of Garudimimus were made in posterior examinations of the specimen, but most of them were criticized during its comprehensive redescription in 2005. Extensive undescribed ornithomimosaur remains at the type locality of Garudimimus may represent additional specimens of the genus.
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Saichania is a genus of herbivorous ankylosaurid dinosaur from the Late Cretaceous period of Mongolia and China.
Varanoidea is a superfamily of lizards, including the well-known family Varanidae. Also included in the Varanoidea are the Lanthanotidae, and the extinct Palaeovaranidae.
A nose is a protuberance in vertebrates that houses the nostrils, or nares, which receive and expel air for respiration alongside the mouth. Behind the nose are the olfactory mucosa and the sinuses. Behind the nasal cavity, air next passes through the pharynx, shared with the digestive system, and then into the rest of the respiratory system. In humans, the nose is located centrally on the face and serves as an alternative respiratory passage especially during suckling for infants. The protruding nose that is completely separate from the mouth part is a characteristic found only in therian mammals. It has been theorized that this unique mammalian nose evolved from the anterior part of the upper jaw of the reptilian-like ancestors (synapsids).
Estesia is an extinct genus of Late Cretaceous anguimorph lizard found in the Gobi Desert in Mongolia.
Saharastega is an extinct genus of basal temnospondyl which lived during the Late Permian period, around 251 to 260 million years ago. Remains of Saharastega, discovered by paleontologist Christian Sidor at the Moradi Formation in Niger, were described briefly in 2005 and more comprehensively in 2006. The description is based on a skull lacking the lower jaws.
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Platynota is a polyphyletic group of anguimorph lizards and thus belongs to the order Squamata of the class Reptilia. Since it was named in 1839, it has included several groups, including monitor lizards, snakes, mosasaurs, and helodermatids. Its taxonomic use still varies, as it is sometimes considered equivalent to the group Varanoidea and other times viewed as a distinct group. It is phylogenetically defined as a clade containing Varanidae. It also includes many extinct species.
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Ovoo gurvel is an extinct varanid lizard from the Late Cretaceous of Mongolia. It is one of the smallest and earliest monitor lizards. It was described in 2008. Ovoo possesses a pair of two small bones in its skull that are not seen in any other lizard.
Aiolosaurus is an extinct genus of monitor lizard from the Late Cretaceous of Mongolia. The type and only species, Aiolosaurus oriens, was named in 2000 from Ukhaa Tolgod, a rich fossil site in the Campanian-age Djadochta Formation.
Cherminotus is an extinct genus of monitor lizard from the Late Cretaceous of Mongolia. The type and only species, Cherminotus longifrons, was named in 1984.
Telmasaurus is an extinct genus of varanoid lizard from the Late Cretaceous of Mongolia. Fossils have been found from the Djadokha and Barun Goyot Formations that date between the early and middle Campanian stage from approximately 80 to 75 million years ago. The type species Telmasaurus grangeri was named in 1943.
Hoburogekko is an extinct genus of gecko that includes a single species, Hoburogekko suchanovi, from the Early Cretaceous of Mongolia. It is known from two fossil specimens, one preserving the front part of the skull and the other preserving part of the lower jaw. Hoburogekko is one of four known Mesozoic geckos or gecko-like lizards, the others being Cretaceogekko from the Early Cretaceous of Burma, AMNH FR21444, an undescribed specimen from a slightly older deposit in Mongolia, and Gobekko from the Late Cretaceous of Mongolia. Hoburogekko is the third oldest known gecko behind AMNH FR21444 and Cretaceogekko.
Myrmecodaptria is an extinct genus of scleroglossan lizard from the Late Cretaceous Djadokhta Formation in Ukhaa Tolgod, Mongolia. The type and only species, Myrmecodaptria microphagosa, was named in 2000 by paleontologists Gao Keqin and Mark Norell. Myrmecodaptria is known from a single holotype skull and lower jaws. It is distinguished from all other lizards by its extremely elongated skull. The eyes are placed close to the snout, which is short and rounded. The top of the skull is covered in bony knobs called osteoderms. The parietal bone at the back of the skull is elongated and about as long as the frontal bones, which are the usually the longest bones along the top of the skull in lizards. The squamosal bone at the back of the skull reaches forward to connect with the jugal bone behind the eye, forming a thin arch between the temporal fenestrae. Myrmecodaptria also has fewer and more widely spaced teeth in its jaws than do most other lizards.
Gobivenator is an extinct genus of troodontid theropod dinosaur known from the late Campanian Djadokhta Formation of central Gobi Desert, Mongolia. It contains a single species, Gobivenator mongoliensis. G. mongoliensis is known from a single individual, which represents the most complete specimen of a Late Cretaceous troodontid currently known.
Polrussia is an extinct genus of iguanian lizards dating to the Late Cretaceous epoch, found in what is now Mongolia. It belongs to a group of extinct iguanians called Gobiguania that was endemic to the Gobi Desert during the Late Cretaceous. The type species Polrussia mongoliensis was named in 1991 on the basis of a skull found in the Barun Goyot Formation. The genus name refers to the Polish and Russian paleontologists who worked together to find and describe the material. Polrussia has a short skull, slightly pointed and flattened snout, and large eye sockets. The teeth each have one cusp, as opposed to the multiple cusps seen in some other gobiguanians. The skull is only 1.2 centimetres (0.47 in) long, making Polrussia one of the smallest gobiguanians.
Igua is an extinct genus of iguanian lizards belonging to a group called Gobiguania that was endemic to the Gobi Desert during the Late Cretaceous. The type species Igua minuta was named in 1991 on the basis of a skull from the Barun Goyot Formation in Mongolia. The skull itself is very small, only 14 millimetres (0.55 in) long, and may have belonged to a juvenile given that it possesses a large fontanelle and that many of the bones are unfused. The snout-vent length of the individual is estimated to have been 55 to 65 millimetres. Igua differs from related gobiguanians like Polrussia in having a more rounded skull. It is similar in appearance to the living genera Liolaemus and Tropidurus. The teeth are tricuspid and pleurodont, meaning they are attached to inner surfaces of the jaws.
Coeruleodraco is an extinct genus of choristoderan known from the Late Jurassic (Oxfordian) Tiaojishan Formation in China. Coeruleodraco is significant as the most complete Jurassic choristodere taxon, as the only other named Jurassic choristodere Cteniogenys is based on fragmentary remains. Although similar to Philydrosaurus in its proportions and postcranial characters, it is distinct in retaining several apparently plesiomorphic characters, including a short snout, paired external nares and an open lower temporal fenestra.
