Amniotes are tetrapod vertebrate animals belonging to the clade Amniota, a large group that comprises the vast majority of living terrestrial and semiaquatic vertebrates. Amniotes evolved from amphibian ancestors during the Carboniferous period and further diverged into two groups, namely the sauropsids and synapsids, an event that marks the appearance of Amniota, according to the definition established under the PhyloCode. This basal divergence within Amniota has been dated by molecular studies at 310–329 Ma or 312–330 Ma, but the presence of Hylonomus at Joggins implies a minimal age of about 317 Ma. A fossilized birth-death process study of early amniotes suggested an age of 322–340 Ma. Amniotes are distinguished from the other living tetrapod clade — the non-amniote lissamphibians — by the development of three extraembryonic membranes, thicker and keratinized skin, and costal respiration. Additional unique features are the presence of adrenocortical and chromaffin tissues as a discrete pair of glands near their kidneys, which are more complex, the presence of an astragalus for better extremity range of motion, the diminished role of skin breathing, and the complete loss of metamorphosis, gills, and lateral lines.
"Labyrinthodontia" is an informal grouping of extinct predatory amphibians which were major components of ecosystems in the late Paleozoic and early Mesozoic eras. Traditionally considered a subclass of the class Amphibia, modern classification systems recognize that labyrinthodonts are not a formal natural group (clade) exclusive of other tetrapods. Instead, they consistute an evolutionary grade, ancestral to living tetrapods such as lissamphibians and amniotes. "Labyrinthodont"-grade vertebrates evolved from lobe-finned fishes in the Devonian, though a formal boundary between fish and amphibian is difficult to define at this point in time.
Lepospondyli is a diverse taxon of early tetrapods. With the exception of one late-surviving lepospondyl from the Late Permian of Morocco, lepospondyls lived from the Visean stage of the Early Carboniferous to the Early Permian and were geographically restricted to what is now Europe and North America. Five major groups of lepospondyls are known: Adelospondyli; Aïstopoda; Lysorophia; Microsauria; and Nectridea. Lepospondyls have a diverse range of body forms and include species with newt-like, eel- or snake-like, and lizard-like forms. Various species were aquatic, semiaquatic, or terrestrial. None were large, and they are assumed to have lived in specialized ecological niches not taken by the more numerous temnospondyl amphibians that coexisted with them in the Paleozoic. Lepospondyli was named in 1888 by Karl Alfred von Zittel, who coined the name to include some tetrapods from the Paleozoic that shared some specific characteristics in the notochord and teeth. Lepospondyls have sometimes been considered to be either related or ancestral to modern amphibians or to Amniota. It has been suggested that the grouping is polyphyletic, with aïstopods being primitive stem-tetrapods, while recumbirostran microsaurs are primitive reptiles.
Reptiliomorpha is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes Homo sapiens, but not Ascaphus truei. Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing Homo sapiens, but not Pipa pipa, Caecilia tentaculata, and Siren lacertina.
Anthracosauria is an order of extinct reptile-like amphibians that flourished during the Carboniferous and early Permian periods, although precisely which species are included depends on one's definition of the taxon. "Anthracosauria" is sometimes used to refer to all tetrapods more closely related to amniotes such as reptiles, mammals, and birds, than to lissamphibians such as frogs and salamanders. An equivalent term to this definition would be Reptiliomorpha. Anthracosauria has also been used to refer to a smaller group of large, crocodilian-like aquatic tetrapods also known as embolomeres.
Diadectes is an extinct genus of large reptiliomorphs or synapsids that lived during the early Permian period. Diadectes was one of the first herbivorous tetrapods, and also one of the first fully terrestrial vertebrates to attain large size.
Diadectomorpha is a clade of large tetrapods that lived in Euramerica during the Carboniferous and Early Permian periods and in Asia during Late Permian (Wuchiapingian), They have typically been classified as advanced reptiliomorphs positioned close to, but outside of the clade Amniota, though some recent research has recovered them as the sister group to the traditional Synapsida within Amniota, based on inner ear anatomy and cladistic analyses. They include both large carnivorous and even larger herbivorous forms, some semi-aquatic and others fully terrestrial. The diadectomorphs seem to have originated during late Mississippian times, although they only became common after the Carboniferous rainforest collapse and flourished during the Late Pennsylvanian and Early Permian periods.
Seymouria is an extinct genus of seymouriamorph from the Early Permian of North America and Europe. Although they were amphibians, Seymouria were well-adapted to life on land, with many reptilian features—so many, in fact, that Seymouria was first thought to be a primitive reptile. It is primarily known from two species, Seymouria baylorensis and Seymouria sanjuanensis. The type species, S. baylorensis, is more robust and specialized, though its fossils have only been found in Texas. On the other hand, S. sanjuanensis is more abundant and widespread. This smaller species is known from multiple well-preserved fossils, including a block of six skeletons found in the Cutler Formation of New Mexico, and a pair of fully grown skeletons from the Tambach Formation of Germany, which were fossilized lying next to each other.
Casineria is an extinct genus of tetrapodomorph which lived about 340–334 million years ago in the Mississippian epoch of the Carboniferous period. Its generic name, Casineria, is a latinization of Cheese Bay, the site near Edinburgh, Scotland, where the holotype fossil was found. When originally described in 1999, it was identified as a transitional fossil noted for its mix of basal (amphibian-like) and advanced (reptile-like) characteristics, putting it at or very near the origin of the amniotes, the group containing all mammals, birds, modern reptiles, and other descendants of their reptile-like common ancestor. However, the sole known fossil is lacking key elements such as a skull, making exact analysis difficult. As a result, the classification of Casineria has been more controversial in analyses conducted since 1999. Other proposed affinities include a placement among the lepospondyls, seymouriamorphs, "gephyrostegids", or as a synonym of Caerorhachis, another controversial tetrapod which may have been an early temnospondyl.
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous (Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous (Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape.
Discosauriscus was a small seymouriamorph which lived in what is now Central and Western Europe during the latest Carboniferous and in the Early Permian Period. Its best fossils have been found in the Broumov and Bačov Formations of Boskovice Furrow, in the Czech Republic.
Chroniosuchus is an extinct genus of chroniosuchid from the upper Permian period. The genus was first named by Vjuschkov in 1957.
Gephyrostegus is a genus of extinct gephyrostegid reptiliomorph amphibian. It was a small animal at 22 cm snout-vent length, of generally lizard-like build and presumably habit. It had large eyes and a large number of small, pointed teeth, indicating it was an active insectivorous hunter. The remains have been found in Nýřany, Czech Republic, dating from around 310 million years ago.
Kotlassia extinct genus of kotlassiine seymouriamorph from the Late Permian of Russia. The type, and currently only, species is K. prima.
Karpinskiosaurus is an extinct genus of seymouriamorphs. It includes two species: Karpinskiosaurus secundus and Karpinskiosaurus ultimus. Karpinskiosaurus secundus is represented by two specimens with skull lengths of about 75 mm. All specimens of K. ultimus are smaller than those of K. secundus. Total length of the reptiliomorph was about 50–75 cm.
Ariekanerpeton is an extinct genus of seymouriamorph from the lower Permian. Fossils have been found from Tajikistan representing over 900 individuals of various stages of ontogenic development. However, it is thought that none of these specimens are of fully mature animals as poor bone ossification is present and the neural arches are paired and disarticulated from the pleurocentra.
Utegenia is a genus of early tetrapod. It is usually regarded as a basal seymouriamorph, but sometimes included in the Discosauriscidae or as a sister taxon of the latter. Only one species, Utegenia shpinari, found from Kazakhstan, is known. Urumqia, another basal seymouriamorph, from Ürümqi, Xinjiang of China is probably a junior synonym of Utegenia.
Makowskia is an extinct genus of discosauriscid seymouriamorph known from the early Permian of Boskovice Furrow, in the Czech Republic. It was first named by Jozef Klembara in 2005 and the type species is Makowskia laticephala. The generic name honors Alexander Makowsky for describing the first specimens of discosauriscids from the Boskovice Furrow, and the specific name means “broad” + “head”. Makowskia is known only from one specimen, the holotype SNMZ 26506, a skull and anterior portion of postcranial skeleton. A phylogenetic analysis places Makowskia as the sister taxon to Spinarerpeton.
The Organ Rock Formation or Organ Rock Shale is a formation within the late Pennsylvanian to early Permian Cutler Group and is deposited across southeastern Utah, northwestern New Mexico, and northeastern Arizona. This formation notably outcrops around Canyonlands National Park, Natural Bridges National Monument, and Monument Valley of northeast Arizona, southern Utah. The age of the Organ Rock is constrained to the latter half of the Cisuralian epoch by age dates from overlying and underlying formations. Important early terrestrial vertebrate fossils have been recovered from this formation in northern Arizona, southern Utah, and northern New Mexico. These include the iconic Permian terrestrial fauna: Seymouria, Diadectes, Ophiacodon, and Dimetrodon. The fossil assemblage present suggests arid environmental conditions. This is corroborated with paleoclimate data indicative of global drying throughout the early Permian.
This list of fossil amphibians described in 2020 is a list of new taxa of fossil amphibians that were described during the year 2020, as well as other significant discoveries and events related to amphibian paleontology that occurred in 2020.
